Form and Function: A Contribution to the History of Animal Morphology — John Shaqi
Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
The principle of the adaptedness of parts may be used as an
explanatory principle, enabling the naturalist to trace out in detail
the interdependence of functions and their organs. When you have
discovered how one organ is adapted to another and to the whole, you
have gone a certain way towards understanding it. That is using
teleology as a regulative principle, in Kant's sense of the word.
Cuvier was indeed a teleologist after the fashion of Kant, and there
can be no doubt that he was influenced, at least in the exposition of
his ideas, by Kant's _Kritik der Urtheilskraft_, which appeared ten
years before the publication of the _Lecons d'Anatomie Comparee_.
Teleology in Kant's sense is and will always be a necessary postulate
of biology. It does not supply an explanation of organic forms and
activities, but without it one cannot even begin to understand living
things. Adaptedness is the most general fact of life, and innumerable
lesser facts can be grouped as particular cases of it, can be, so far,
understood.
Cuvier's famous principle of correlation, the corner-stone of his
work, is simply the practical application to the facts of structure of
the principle of functional adaptedness. By the principle of
correlation, from one part of an animal, given sufficient knowledge of
the structure of its like, you can in a general way construct the
whole. "This must necessarily be so: for all the organs of an animal
form a single system, the parts of which hang together, and act and
re-act upon one another; and no modifications can appear in one part
without bringing about corresponding modifications in all the
rest."[47] The logical basis of the principle is sound. The functions
of the parts are all intimately bound up with one another, and one
function cannot vary without bringing in its train corresponding
modifications in the others. Structure and function are bound up
together; every modification of a function entails therefore the
modification of an organ. Hence from the shape of one organ you can
infer the shape of the other organs--if you have sufficiently
extensive empirical knowledge of functions, and of the relation of
structure to function in each kind of organ. Given an alimentary canal
capable of digesting only flesh, and possessing therefore a certain
form, you know that the other functions must be adapted to this
particular function of the alimentary canal. The animal must have keen
sight, fine smell, speed, agility, and strength in paws and jaws.
These particular functions must have correspondingly modified organs,
well-developed eyes and ears, claws and teeth. Further, you know from
experience that such and such definitely modified organs are
invariably found with the carnivorous habit, carnassial teeth, for
example, and reduced clavicles. From a "carnivorous" alimentary canal,
then, you can infer with certainty that the animal possessed
carnassial teeth and the other structural peculiarities of carnivorous
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