Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
If the same materials of organisation are present in all animals, and if
they are arranged always in the same positions relatively to one
another, how does it come about that animal forms are so varied, what
explanation can be offered of the diversities of organic structure?
Geoffroy's main answer to this question is his _Loi de balancement_. The
law was enunciated by him already in 1807.[114] We take the following
quotation, which represents his thought most nearly, from the _Cours de
l'histoire naturelle des Mammiferes_ (1829). "According to our manner of
regarding the organisation of mammals, there is only a single animal
modified by the inverse reciprocal variation of all or some of its
parts. Now, from the fact that there is only one single general animal,
it follows that for each section of its components or for each of its
organs there is available only a given quantity of formative materials.
Now suppose that the distribution of these materials has not been made
in such a way as to ensure an exact equilibrium between all the parts
concerned, one organ will get more than its share, another less. My law
of the compensation of organs is founded on these principles" (i.,
_Lecon_ 16, p. 12). "The atrophy of one organ turns to the profit of
another; and the reason why this cannot be otherwise is simple, it is
because there is not an unlimited supply of the substance required for
each special purpose."[115] The nutritive material available is limited
for each species; if one part gets more than its share the other parts
must get less--that is all the law means. As an example, take the
minuteness of the episternals and xiphisternals in birds, as contrasted
with the huge size of the entosternal. "The minuteness of the
episternals and xiphisternals might be imputed to this gigantic piece
diverting to its own profit the nutritive fluid, since the bigger it is
the smaller these are."[116]
One has constantly to remember in dealing with Geoffroy's theories that
he was not an evolutionist, but purely a morphologist. It is therefore,
perhaps, to ask too much to require of him an explanation of the causes
of diversity. The morphologist describes, classifies, generalises; he
does not seek for causes. But we must leave this question aside in order
to discuss how far Geoffroy's theory of the unity of plan and
composition fits the facts. As Geoffroy himself admitted on several
occasions, his theory was an _a priori_ one, a theory hit upon by hasty
induction, then erected into a principle and imposed upon the facts. No
more than Goethe did he extract his principle from a sufficient mass of
data.
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