Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.Scudder, Samuel Hubbard
Science
Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.
Scudder, Samuel Hubbard
Butterflies, Fossil
“In order to show the extent to which the Jermyn street example is
deficient, I have restored it (fig. 5 [our fig. 7]), filling in the
blanks from Mr. Charlesworth’s specimens. By comparing the latter with
the wing of _Dasyophthalma_ (fig. 1), and _Cicada_ (fig. 2), one may come
to a pretty accurate conclusion as to the group of insects to which it
ought to be referred.”[BC]
The neuration of Lepidoptera as a group is the simplest in the whole
order of insects, if we except that of the elytra of Coleoptera; this
is due, doubtless, to the fact that their wings are heavily scaled,
concealing the nervures; just as in Coleoptera, the thickness and opacity
of the fore wings often completely masks the neuration.
[Illustration: Fig. 7.
Palæontina oolitica Butl. The neuration, after Butler’s second sketch.]
The normal number of veins in the wings of insects is six, disposed to
a certain extent in pairs; the middle pair usually ramify to a greater
extent than the others, and support most of the membrane of the wing.
In butterflies the foremost vein is always absent and very commonly the
hindmost, so that there are but five (often but four) principal veins,
usually designated, though not very appropriately: costal, subcostal,
median, submedian and (when present) internal, reciting them in their
order from in front backward. The costal, submedian and internal nervures
are invariably simple and terminate at the margin, or are occasionally
lost in the membrane of the wing. The subcostal and median nervures,
on the other hand, are as invariably forked, and with their branches
support nearly the entire wing; the subcostal nervure curves downward
and the median upward so as to meet, or nearly to meet, not far from the
middle of the wing, and to enclose between them a large space called
the discoidal cell; the branches of the median nervure are all thrown
off from its lower edge before union with the subcostal; the principal
branches of the subcostal nervure are, on their side, thrown off from
its upper edge; but, as the nervure curves downward at the extremity of
the cell, another set is thrown off (at least in the fore wings) from
the lower edge; and it is these veins, rather than the subcostal nervure
proper, which unite with the median to close the cell.[BD] None of the
median, nor any of the inferior subcostal nervules are ever branched;
but at the apex of the wing, where the play of neuration is usually the
greatest, the last superior subcostal nervule is occasionally forked in
the front wing. This is the only forked branchlet in either of the wings.
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