Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.Scudder, Samuel Hubbard
Science
Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.
Scudder, Samuel Hubbard
Butterflies, Fossil
The last figure of _P. oolitica_ given by Mr. Butler agrees in all its
essential features with his first illustration. They both represent
a front wing with four principal nervures,—costal, subcostal, median
and submedian; the costal nervure is swollen at the base and extends,
unbranched, to the tip of the wing; the median nervure is three-branched,
the three forks simple, equidistant, emitted from the apical half of
the vein, which at its extremity is united by a cross vein to a branch
of the subcostal, closing the cell; the submedian nervure is simple and
divides the space between the median vein and the margin of the wing.
So far all is in accordance with the lepidopterous type; but when we
examine the subcostal vein, which occupies nearly half the wing, the
resemblance ceases altogether. This vein is represented as bearing no
superior branches, but as sending out from its _inferior_ surface three
distinct veinlets, the first and second of which again emit a tributary
from their inferior surfaces. This is a structural anomaly which finds
no counterpart whatsoever in any family of butterflies. So that should
we accept Mr. Butler’s own sketch of the fossil as correct, it would be
impossible to consider the wing that of a butterfly.
In his description of the insect Mr. Butler compares the neuration to
that of Caligo, and says its nearest allies are Caligo, Dasyophthalma and
Brassolis. In his latter paper he figures the wing of a Dasyophthalma by
way of comparison. In the genera named all the branches of the subcostal
nervure are simple, and are thrown off from the superior surface,
excepting the single set which is emitted from beneath, and which marks
(as in all butterflies) the limit of the discoidal cell; this corresponds
fairly with the first set of inferior veins emitted by the subcostal vein
in the fossil; for the other sets, however, no counterpart will be found
in the living types.
[Illustration: Fig. 8.
Palæontina oolitica Butl. Corrected sketch of the neuration.]
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