Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.Scudder, Samuel Hubbard
Science
Fossil Butterflies: Memoirs of the American Association for the Advancement of Science, I.
Scudder, Samuel Hubbard
Butterflies, Fossil
It was probably Mr. Butler’s want of familiarity with fossils that led
him to overlook several features which can be seen in these originals.
Having first traced the outline of the wing and the general course of
the veins directly from the specimens, I subsequently filled in by
measurement all the other parts which I could follow, studying each vein,
or supposed vein, with the utmost care, from one end to the other of its
course. The result of that study is presented in fig. 8, which differs
essentially in its details from the illustrations given by Butler, and
looks, as he himself confesses, “exceedingly anti-lepidopterous.” In the
first place, the wing is much narrower than depicted by him; and at the
extremity of a vein (the submedian vein of Butler’s sketch) there is a
slight but decided bending inward of the membrane, as very frequently
occurs at the line of demarcation between the middle and inner area
of the wing in all or nearly all the lower suborders of insects, but
never, so far as I am aware, in Lepidoptera. What he has given as a
simple costal vein is neither swollen at the base nor simple, but has
two inferior branches near the middle of the wing, united near their
origin by an oblique cross vein. Branching of the costal vein is unknown
in Lepidoptera; but if it should be claimed that this might be the
subcostal, just as much difficulty will be encountered with the structure
and relationship of the veinlets below, which must then be considered
as belonging to the median vein; in no Lepidoptera can any such
irregularity be shown, nor so disproportionate a magnitude of the area
covered by the median nervure and its branches; a branched internal vein
and cross-veins, which probably united all the longitudinal nervures at
no great distance from the outer border (but which can only be certainly
predicated for the lower three median interspaces), place this insect
wholly beyond the pale of the Lepidoptera. It is but fair to say that
Mr. Butler, having examined the original after he had in his possession
a tracing of fig. 8, denies the existence of the cross-veins; there is
one point, however, which an unprejudiced examination of the fossil
cannot fail to show; that Butler’s “fourth branch” of the subcostal[BE]
arises not from his third branch, but from his upper discoidal vein;
if he can reconcile either this or the points already referred to (on
the supposition that his sketch is otherwise an accurate one) with the
neuration of any group of butterflies, the writer will be the first to
acknowledge it.
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