Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In the present case I am disposed to regard the _Caulerpa_-like casts
as of animal rather than plant origin. The clavate branches have
the form of very deep moulds in the hard brown rock which have been
filled in with blue mud. It is hardly conceivable that the branches
of a soft watery plant such as _Caulerpa_ could leave more than a
faint impression on an old sea-floor. The specimens occur in different
positions in the matrix of the rock and they are not confined to
the lines of bedding; in none of the examples is there any trace of
carbonaceous matter in association with the deep moulds. On the whole,
then, this Kimeridge fossil cannot, I believe, be accepted as an
authentic example of a Mesozoic _Caulerpa_.
It is not improbable that some of the supposed fossil algae may be
casts of egg-cases or spawn-clusters of animals. In Ellis’ Natural
History of the Corallines[278] there is a drawing representing a number
of disc-like ovaries attached to a tough ligament, and referred to the
mollusc _Buccinum_, which bears a certain resemblance to the Weymouth
fossil. A similar body is figured by Fuchs[279] in an important memoir
on supposed fossil algae.
It is not suggested that the _Caulerpa Carruthersi_ of Murray should be
regarded as the cast of some molluscan egg-case attached to a slender
axis, but it is important to bear in mind the possibility of matching
such extremely doubtful fossils with other organic bodies than the
thallus of a _Caulerpa_. In an example of an egg-case in the Cambridge
Zoological Museum, referred to a species of _Pyrula_, there is a hard,
long and slender axis, bearing a series of semicircular chambers
divided into radial compartments. The whole is hard and horny and might
well be preserved as a fossil.
β. Codiaceae.
The members of this Order present a considerable diversity of form as
regards the shape of the plant-body; the thallus of some species is
encrusted with carbonate of lime. The order is widely distributed in
tropical and temperate seas.
Among the recent genera _Penicillus_ and _Codium_ may be chosen as
important types from the point of view of fossil representatives.
_Codium._
The thallus of _Codium_ consists of a spongy mass of tubular
cell-branches which are differentiated into two fairly distinct
regions, an outer peripheral layer in which the branches have long
club-shaped terminations, and an inner region consisting of loosely
interwoven filaments.
_Codium Bursa_ L. and _C. tomentosum_ Huds. are two well-known British
species, the former presents the appearance of a spongy ball of
cells, and in the latter the thallus is divided up into dichotomously
forked branches[280]. In this genus the thallus is not encrusted with
carbonate of lime, at least in recent species.
_Sphaerocodium._ Fig. 37, D.
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