Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Rothpletz[281] instituted this genus for certain small spherical
or oval bodies varying from 1 mm. to 2 cm. in diameter, which have
been found on crinoid stems or shell fragments of Triassic age. Each
spherical body consists of dichotomously branched single-celled
filaments, between 50 and 100µ in breadth, and from 300–500µ in height.
The tubular cavities occasionally swell out into spherical spaces which
are regarded by Rothpletz as sporangia.
There is not sufficient evidence that _Sphaerocodium Bornemanni_ Roth.
has been correctly referred to the Codiaceae. The sporangia-like
swellings described by the author of the species are not by any means
conclusive as characters of important taxonomic value. Figure 37, D,
illustrates the general structure of the fossil as seen in a transverse
section of one of the calcareous grains.
Like _Girvanella_, which has been referred by some writers to the
Siphoneae, _Sphaerocodium_ occurs in the form of oolitic grains. In
the Triassic Raibler and St Cassian beds of the Tyrol, as well as in
rocks of Rhaetic age in the Eastern Alps, it makes up large masses
of limestone. Rothpletz compares the structure of this genus with
that of the recent alga _Codium adhaerens_ Ag., but it is wiser to
regard such tubular structures as _Girvanella_, _Siphonema_[282] and
_Sphaerocodium_ as closely allied organisms, which are probably algae,
but too imperfectly known to be referred to any particular family.
_Penicillus._
The recent genus _Penicillus_ is one of those algae formerly included
among animals. Fig. 33, O, has been copied from a drawing of a species
of _Penicillus_ given by Lamouroux[283] under the generic name of
_Nesea_ in his treatise on the genera of Polyps published in 1821. He
describes the genus as a brush-like Polyp with a simple stem.
The thallus consists of a stout stem terminating in a brush-like tuft
of fine dichotomously-branched filaments. The apical branches are
divided by regular constrictions into short oval or rod-like segments
which may be encrusted with carbonate of lime. A few of the segments
from the terminal tuft of a recent _Penicillus_ are shown in fig. 35,
E. Each of these calcareous segments has the form of an oval shell
perforated at each end, and the wall is pierced by numerous fine
canals. _Penicillus_ is represented by about 10 recent species, which
with one exception live in tropical seas.
The recognition of _Penicillus_, or a very similar type, in a fossil
condition is due to Munier-Chalmas[284]. This keen observer has
rendered great service to palaeobotany by directing attention to the
calcareous algae in the Paris basin beds, and by proving that many of
the fossils from these Tertiary deposits have been erroneously included
by previous writers among the Foraminifera[285]. It is greatly to be
desired that Prof. Munier-Chalmas may soon publish a monograph on the
fossil Siphoneous forms of which he possesses a unique collection.
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