Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
1. _Marchantites erectus_ (Leckenby). Fig. 49. This species may be
described as follows: The thalloid body is divided into spreading
dichotomously branched segments, obtusely pointed apically. The
slightly wrinkled surface shows a distinct and comparatively broad
darker and shorter median band, with lighter coloured and thinner
margins.
[Illustration: FIG. 49. _Marchantites erectus_ (Leck.). From the
type-specimen in the Woodwardian Museum. Nat. size.]
In 1864 Leckenby described this plant from the Lower Oolite beds
of the Yorkshire coast near Scarborough, as _Fucoides erectus_,
regarding it as a fossil alga. I recently pointed out that the general
appearance and mode of occurrence of the specimens suggest a liverwort
rather than an alga, and proposed the substitution of the genus
_Marchantites_[460]. It would, however, be unwise to speak with any
great confidence as to the real affinities of the fossil.
The example shown in the figure is the type-specimen of Leckenby[461]:
the breadth of the branches is about 3 mm. Under a low magnifying power
the surface shows distinct and somewhat oblique wrinklings, the general
appearance being very similar to that of some recent forms of the genus
_Marchantia_.
A closely allied species has recently been described from the
Wealden beds of Ecclesbourne, near Hastings, on the Sussex coast, as
_Marchantites Zeilleri_ Sew.[462].
In a recent monograph on Jurassic plants from Poland, apparently
containing much that is of the greatest value, but which is
unfortunately written in the Polish language, Raciborski[463] describes
a new species of thalloid Liverwort under the name of _Paleohepatica
Rostafinski_. The specimens are barren plants larger than any Jurassic
species hitherto described; they agree closely in habit with Saporta’s
Tertiary species _Marchantites Sezannensis_.
2. _Marchantites Sezannensis Saporta._ Fig. 50. The body is broadly
linear and dichotomously branched, with a somewhat undulating margin.
Midrib on the dorsal surface depressed, but more prominent on the
ventral surface. The upper surface is divided into hexagonal areas, in
each of which occurs a central pore. There are two rows of scales along
the median line on the lower surface. Stalked male receptacles.
Brongniart[464] first mentioned this fossil hepatic, which was found in
the calcareous travertine of Sézanne of Oligocene age in the Province
of Marne. The specimens figured by Saporta[465] show very clearly
the characters of one of the _Marchantiaceae_, and in this case we
have the additional evidence of the characteristic male receptacles
which are given off from a point towards the apex of the lobes, and
arise from a slight median depression. In one of Saporta’s figures
(reproduced in fig. 50 _A_) there are represented some median scars
which may mark the position of cups similar to those which occur on
recent species of _Marchantia_, and in which gemmæ or bulbils are
produced.
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