Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The specimens of fossil Equisetums are usually readily recognised by
the coherent leaf-segments in the form of nodal sheaths resembling
those of recent species. The tissues of the cortex and central
cylinder are occasionally represented by a thin layer of coal pressed
on to the surface of a sandstone cast, or covering a flattened
stem-impression on a piece of shale. It is sometimes possible under
the microscope to recognise on the carbonised epidermal tissues
the remains of a surface-ornamentation similar to that in recent
species, which is due to the occurrence of siliceous patches on the
superficial cells. Longitudinal rows of stomata may also be detected
under favourable conditions of preservation. The nodal diaphragms
of stems have occasionally been preserved apart, but such circular
and radially-striated bodies may be misleading if found as isolated
objects. Casts of the wide hollow pith of _Equisetites_, with
longitudinal ridges and grooves, and fairly deep nodal constrictions,
have often been mistaken for the medullary casts of _Calamites_.
Several species of _Equisetites_ have been recorded from the Upper
Coal-Measures and overlying Permian rocks, but these present special
difficulties. In one instance described below, (_Equisetites
Hemingwayi_ Kidst.), the species was founded on a cast of what appeared
to be a strobilus made up of sporophylls similar to those in an
_Equisetum_ cone. In other Permo-Carboniferous species the choice of
the generic name _Equisetites_ has been determined by the occurrence
of leaf-sheaths either isolated or attached to the node of a stem. The
question to consider is, how far may the Equisetum-like leaf-sheath
be regarded as a characteristic feature of _Equisetites_ as distinct
from _Calamites_? In the genus _Calamites_ the leaves are generally
described as simple linear leaves arranged in a whorl at the nodes,
but not coherent in the form of a sheath (fig. 85). The fusion of
the segments into a continuous sheath or collar is regarded as a
distinguishing characteristic of _Equisetites_ and _Equisetum_. The
typical leaf-sheath of a recent Horse-tail has already been described.
In some species we have fairly large and persistent free teeth on the
upper margin of the leaf-sheath, but in other Equisetums the rim of
the sheath is practically straight and has a truncated appearance,
the distal ends of the segments being separated from one another by
very slight depressions, as in a portion of the sheath of _Equisetum
ramosissimum_ Desf. of fig. 58, _C_. In other leaf-sheaths of this
species there are delicate and pointed teeth adherent to the margin
of the coherent segments; the teeth are deciduous, and after they have
fallen the sheath presents a truncated appearance. This difference
between the sheaths to which the teeth are still attached and those
from which they have fallen is illustrated by fig. 58, _B_ and _C_;
it is one which should be borne in mind in the description of fossil
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