Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
species, and has probably been responsible for erroneous specific
diagnoses. In some recent Horse-tails the sheath is occasionally
divided in one or two places by a slit reaching to the base of the
coherent segments[498]; this shows a tendency of the segments towards
the free manner of occurrence which is usually considered a Calamitean
character. In certain fossils referred to the genus _Annularia_, the
nodes bear whorls of long and narrow leaves which are fused basally
into a collar (fig. 58, _D_). There are good grounds for believing that
at least some Annularias were the foliage shoots of true Calamites.
Again, in some species of _Calamitina_, a sub-genus of _Calamites_,
the leaves appear to have been united basally into a narrow sheath.
We see, then, that it is a mistake to attach great importance to the
separate or coherent character of leaf-segments in attempting to draw
a line between the true _Calamites_ and _Equisetites_. Potonié[499]
while pointing out that this distinction does not possess much value
as a generic character, retains the genus _Equisetites_ for certain
Palaeozoic Equisetum-like leaf-sheaths.
[Illustration: FIG. 56. Calamitean leaf-sheath. From a specimen in the
Woodwardian Museum. _a_, base of leaf-sheath; (very slightly
reduced).]
Fig. 56 represents a rather faint impression of a leaf-sheath and
nodal diaphragm. The specimen is from the Coal-Measures of Ardwick,
Manchester. The letter _a_ probably points to the attachment of the
sheath to the node of the stem. The flattened sheath is indistinctly
divided into segments, and at the middle of the free margin there
appears to be a single free tooth. The lower part of the specimen, as
seen in the figure, shows the position of the nodal diaphragm. Between
the diaphragm and the sheath there are several slight ridges converging
towards the nodal line; these agree with the characteristic ridges and
grooves of Calamite casts which are described in detail in Chapter X.
There is another specimen in the British Museum which illustrates,
rather more clearly than that shown in fig. 56, the association of a
fused leaf-sheath with a type of cast usually regarded as belonging
to a Calamitean stem. Some leaf-sheaths of Permian age described by
Zeiller[500] as _Equisetites Vaujolyi_ bear a close resemblance to
the sheath in fig. 58 E. The nature of the true Calamite leaves is
considered more fully on a later page.
[Sidenote: PALAEOZOIC EQUISETITES.]
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