Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Our acquaintance with this species is practically limited to the
casts of stems. A typical stem of _E. columnaris_ measures 3 to 6
cm. in diameter and has fairly long internodes. The largest stem in
the British Museum collection has internodes about 14 cm. long and a
diameter of about 5 cm. In some cases the stem casts show irregular
lateral projections in the neighbourhood of a node, but there is no
evidence that the aerial shoots of this species gave off verticils of
branches. In habit _E. columnaris_ probably closely resembled such
recent species as _Equisetum hiemale_ L., _E. trachyodon_ A. Br. and
others.
The stems often show a distinct swelling at the nodes; this may be due,
at least in part, to the existence of transverse nodal diaphragms which
enabled the dead shoots to resist contraction in the region of the
nodes. The leaf-sheaths consist of numerous long and narrow segments
often truncated distally, as in fig. 58, _B_, and as in the sheath of
such a recent Horse-tail as _E. ramosissimum_ shown in fig. 58, _C_. In
some specimens one occasionally finds indications of delicate acuminate
teeth extending above the limits of a truncated sheath. Brongniart
speaks of the existence of caducous acuminate teeth in his diagnosis of
the species, and the example represented in fig. 58, _B_, demonstrates
the existence of such deciduous appendages. There is a very close
resemblance between the fossil sheath of fig. 58, _B_, with and without
the teeth, and the leaf-sheath of the recent _Equisetum_ in fig. 58,
_C_. In some specimens of _E. columnaris_ in which the cast is covered
with a carbonaceous film, each segment in a leaf-sheath is seen to be
slightly depressed in the median portion, which is often distinctly
marked by numerous small dots, the edges of the segment being flat and
smooth. The median region is that in which the stomata are found and on
which deposits of silica occur.
6. _Equisetites Beani_ (Bunb.). Figs. 60–62.
Bunbury[518] proposed the name _Calamites Beani_ for some fossil stems
from the Lower Oolite beds of the Yorkshire coast, which Bean had
previously referred to in unpublished notes as _C. giganteus_. The
latter name was not adopted by Bunbury on account of the possible
confusion between this species and the Palaeozoic species _Calamites
gigas_ Brongn. The generic name _Calamites_ must be replaced by
_Equisetites_ now that we are familiar with more perfect specimens
which demonstrate the Equisetean characters of the plant.
[Illustration: FIG. 60. _Equisetites Beani_ (Bunb.). ⅔ nat. size.
[After Starkie Gardner (86) Pl. IX. fig. 2.]]
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