Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In the museums of Whitby and Scarborough there are some long specimens,
in one case 44 cm. in length, and 33 cm. in circumference, which
are probably casts of the broad pith-cavity. These casts are often
transversely broken across at the nodes, so that they consist of three
or four separate pieces which fit together by clean-cut faces. This
manner of occurrence is most probably due to the existence of large and
resistant nodal diaphragms which separated the sand-casts of adjacent
internodes. In the York museum there are some large diaphragms, 10
cm. in diameter, preserved separately in a piece of rock containing
a cast of _Equisetites Beani_. The nodal diaphragms of some of the
Carboniferous Calamites were the seat of cork development[523], and
it may be that the frequent preservation of Equisetaceous diaphragms
in Triassic and Jurassic rocks is due to the protection afforded by a
corky investment.
The stem shown in fig. 62 appears to be a portion of a shoot of _E.
Beani_ not far from its apical region. From the lower nodes there
extend clearly marked and regular lines or slight grooves tapering
gradually towards the next higher node; these are no doubt the
impressions of segments of leaf-sheaths. The sheaths themselves have
been detached and only their impressions remain. The flattened bands at
the node of the stem in fig. 60, and shown also in fig. 61, mark the
place of attachment of the leaf-sheaths. On some of these nodal bands
one is able to recognise small scars which are most likely the casts of
outgoing leaf-trace bundles.
Some of the internal casts of this species are marked by numerous
closely arranged longitudinal lines, which are probably the impressions
of the inner face of a central woody cylinder. In the smaller specimen
shown in fig. 62 we have the apical portion of a shoot in which the
uppermost internodes are in an unexpanded condition.
[Illustration: FIG. 62. _Equisetites Beani_ (Bunb.). From a specimen in
the Scarborough Museum. Very slightly reduced.]
It is impossible to give a satisfactory diagnosis of this species
without better material. The plant is characterised chiefly by the
great breadth of the stem, and by the possession of leaf-sheaths
consisting of numerous long and narrow segments. _Equisetites Beani_
must have almost equalled in size the Triassic species, _E. arenaceus_,
described above.
7. _Equisetites lateralis_ Phill. Figs. 58, _F_, 63, and 64.
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