Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The circular depressions a short distance above each node are the
‘branch scars’ of various writers. Schimper suggested that these
radially marked circles might be displaced nodal diaphragms.
Andrae[530] figured the same objects in 1853 but regarded them as
branch scars, although in the specimen he describes, there are several
of them lying apart from the stems, and to one of them is attached
a portion of a leaf-sheath. Solms-Laubach[531] points out that the
internodal position of these supposed scars is an obvious difficulty;
we should not expect to find branches arising from an internode. After
referring to some specimens in the Oxford museum, he adds—“In presence
of these facts the usual explanation of these structures appears to
me, as to Heer, very doubtful.... We are driven to the very arbitrary
assumption that they represent the lowest nodes of the lateral
branches which were inserted above the line of the nodes of the stem.”
Circular discs similar to those of _E. lateralis_ have been found in
the Jurassic rocks of Siberia[532] and elsewhere. There are one or
two examples of such discs from Siberia in the British Museum. If the
nodal diaphragms were fairly hard and stout, it is easy to conceive
that they might have been pressed out of their original position when
the stems were flattened in the process of fossilisation. It is not
quite clear what the radial spoke-like lines of the discs are due to;
possibly they mark the position of bands of more resistant tissue or
of outgoing strands of vascular bundles. A detached diaphragm is seen
in fig. 64 C; in the centre it consists of a flat plate of tissue, and
the peripheral region is traversed by the radiating lines. In the stem
of fig. 64, A the deeply divided leaf-sheaths are clearly seen, and an
imperfect impression of a diaphragm is preserved on the face of the
middle internode. In fig. 64 B a flattened leaf-sheath is shown with
the free acuminate teeth fused basally into a continuous collar[533].
The short piece of stem of _Equisetites lateralis_ shown in fig. 58,
_F_, shows how the free teeth may be outspread in a manner which bears
some resemblance to the leaves of _Phyllotheca_, but a comparison with
the specimens already described, and a careful examination of this
specimen itself, demonstrate the generic identity of the species with
_Equisetites_. The carbonaceous film on the surface of such stems as
those of fig. 58, F, and 64, A, shows a characteristic shagreen texture
which may possibly be due to the presence of silica in the epidermis as
in recent Horse-tails.
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