Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
On the broader stems, such as that of fig. 67, we have clearly marked
narrow grooves and broader and slightly convex ridges, which present
an appearance identical with that of some Calamitean stems. In the
specimen figured by Bunbury[568] in his Pl. X, fig. 6, there is a
circular depression on the line of the node which represents the
impression of the basal end of a branch; on the edges of the node
there are indications of two other lateral branches. The nature of
this stem-cast points umnistakeably to a woody stem like that of
_Calamites_. The precise meaning of the ridges and grooves on the cast
is described in the Chapter dealing with Calamitean plants.
[Sidenote: CALAMOCLADUS.]
Grand’Eury[569] in his monograph on the coal-basin of Gard, has
recently described under the name of _Calamocladus frondosus_ what he
believes to be the leaf-bearing axes of a Calamitean plant. The thicker
branches are almost exactly identical in appearance with the broader
specimens of _Phyllotheca_. The finer branches of _Calamocladus_
bear cup-like leaf-sheaths which are divided into long and narrow
recurved segments (fig. 67, B), precisely as in _Phyllotheca_. These
comparisons lead one to the opinion that the _Phyllotheca_ of Australia
and India may be a close ally of the Permo-Carboniferous Calamitean
plants. The form of the leaf-whorls of _Annularia_ (Calamarian
leaf-bearing branches) and of _Calamocladus_ is of the same type as
in _Phyllotheca_; the character of the medullary casts is also the
same. The nature of the fertile shoot of _Phyllotheca_ described by
Schmalhausen from Siberia, with its alternating whorls of sterile and
fertile leaves, is another point of agreement between this genus and
Calamitean plants. An Equisetaceous species has been described from
the Newcastle Coal-Measures of Australia by Etheridge[570] in which
there are two forms of leaves, some of which closely resemble those of
_Phyllotheca indica_, while others are compared with the sterile bracts
of _Cingularia_, a Calamitean genus instituted by Weiss[571].
When we turn to other recorded forms of _Phyllotheca_ many of
them appear on examination to have been placed in this genus on
unsatisfactory grounds. Heer figures several stem fragments from the
Jurassic rocks of Siberia as _P. Sibirica_ Heer[572], and it was the
resemblance between this form and the English _Equisetites lateralis_
which led to the substitution of _Phyllotheca_ for _Equisetites_ in
the latter species. Without examining Heer’s material it is impossible
to criticise his conclusions with any completeness, but several of
his specimens, appear to possess leaf-sheaths more like those of
_Equisetum_ than of _Phyllotheca_.
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