Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The frequent occurrence of isolated diaphragms and the comparatively
long acuminate teeth of the leaf-sheath afford obvious points of
resemblance to _Equisetites lateralis_. Some of the examples figured
by Heer appear to be stem fragments, with numerous long and narrow
filiform leaves different in appearance from those of other specimens
which he figures. It may be that some of the less distinct pieces
of stems are badly torn specimens in which the internodes have been
divided into filiform threads. Heer also figures a fertile axis
associated with the sterile stems, and this does not, as Heer admits,
show the alternating sterile bracts such as Schmalhausen has described.
So far as it is possible to judge from an examination of Heer’s figures
and a few specimens from Siberia in the British Museum—and this is by
no means a safe basis on which to found definite opinions—there appears
to be little evidence in favour of separating the fossils described
as _Phyllotheca Sibirica_ from _Equisetites_. This Siberian form may
indeed be specifically identical with _Equisetites lateralis_ Phill.
Various species of _Phyllotheca_ have been described from Jurassic and
Upper Palaeozoic rocks in Australia. Some of these possess cup-like
leaf-sheaths, and in the case of the thicker specimens they show
continuous ridges and grooves on the internodes, as well as a habit
of branching similar to that in some of the Italian Phyllothecas. In
some of the stems it is however difficult to recognise any characters
which justify the use of the term _Phyllotheca_. A fragment figured by
Tenison-Woods[573] as a new species of _Phyllotheca_, _P. carnosa_,
from Ipswich, Queensland, affords an example of the worthless material
on which species have not infrequently been founded. The author of
the species describes his single specimen as a “faint impression”;
the figure accompanying his description suggests a fragment of some
coniferous branch, as Feistmantel has pointed out in his monograph on
Australian plants.
It is important that a thorough comparative examination should be
made of the various fossil Phyllothecas with a view to determine
their scientific value, and to discover how far the separation of
_Phyllotheca_ and _Equisetites_ is legitimate in each case. There is
too often a tendency to allow geographical distribution to decide the
adoption of a particular generic name, and this seems to have been
especially the case as regards several Mesozoic and Palaeozoic Southern
Hemisphere plants.
The geological and geographical range of _Phyllotheca_ is a question of
considerable interest, but as already pointed out it is desirable to
carefully examine the various records of the genus before attempting
to generalise as to the range of the species. _Phyllotheca_ is often
spoken of as a characteristic member of the _Glossopteris_ Flora of the
Southern Hemisphere, and its geological age is usually considered to be
Mesozoic rather than Palaeozoic.
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