Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
An important contribution to our knowledge of _Calamites_ was supplied
by Petzholdt[593] in 1841. His main contention was the Equisetaceous
character of this Palaeozoic genus. The external resemblance between
Calamite casts and _Equisetum_ stems had long been recognised, but
after Cotta’s account of the internal structure it was believed that
the apparent relation between _Equisetum_ and _Calamites_ was not
confirmed by the facts of anatomy. Petzholdt based his conclusions
on certain partially preserved Permian stems from Plauenscher Grund,
near Dresden. Although his account of the fossils is not accurate his
general conclusions are correct. The specimens described by Petzholdt
differ from the common Calamite casts in having some carbonised
remnants of cortical and woody tissue. A transverse section of one of
the Plauenscher Grund fossils is shown in fig. 70. The irregular black
patches were described by Petzholdt as portions of cortical tissue,
while he regarded the spaces as marking the position of canals like the
vallecular canals in an _Equisetum_. Our more complete knowledge of the
structure of a Calamite stem enables us to correlate the patches in
which no tissue has been preserved with the broad medullary rays, which
separated the wedge-shaped groups of xylem elements; the latter being
more resistant were converted into a black coaly substance, while the
cells of the medullary rays left little or no trace in the sandstone
matrix. The thin black line, which forms the limit of the drawing in
fig. 70, external to the carbonised wood, no doubt marks the limit
of the cortex, and the appendage indicated in the lower part of the
figure may possibly be an adventitious root. It is interesting to note
that Unger[594] in 1844 expressed the opinion, which we now know to be
correct, that the coaly mass in the specimens described by Petzholdt
represented the wood, and that there was no proof of the existence of
canals in the cortex as Petzholdt believed.
[Illustration: FIG. 70. Transverse section of a Calamite stem, showing
carbonised remnants of secondary wood. From a specimen (no. 40934),
presented to the British Museum by Dr Petzholdt from Plauenscher
Grund, Dresden. ½ nat. size.]
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