Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Turning to Brongniart’s later work[595] we find an important
proposal which led to no little controversy. While retaining the
genus _Calamites_ for such specimens as possess a thin bark and a
ribbed external surface, showing occasional branch-scars at the
nodes, and having such characters as warrant their inclusion in the
Equisetaceae, he proposes a second generic name for other specimens
which had hitherto been included in _Calamites_. The fossils assigned
to his new genus _Calamodendron_ are described as having a thick woody
stem, and as differing from _Equisetum_ in their arborescent nature.
Brongniart’s genus _Calamodendron_ is made to include the plants for
which Cotta instituted the name _Calamitea_, and it is placed among
the Gymnosperms. This distinction between the Vascular Cryptogam
_Calamites_ and the supposed Gymnosperm _Calamodendron_ is based on the
presence of secondary wood in the latter type of stem. The prominence
formerly assigned to the power of secondary thickening possessed by a
plant as a taxonomic feature, is now known to have been the result of
imperfect knowledge. The occurrence of a cambium layer and the ability
of a plant to increase in girth by the activity of a definite meristem,
is a feature which some recent Vascular Cryptogams[596] share with the
higher plants; and in former ages many of the Pteridophytes possessed
this method of growth in a striking degree.
Although Brongniart’s distinction between _Calamites_ and
_Calamodendron_ has not been borne out by subsequent researches, the
latter term is still used as a convenient designation for a special
type of Calamitean structure. One of the earliest accounts of the
anatomy of _Calamodendron_ stems is by Mougeot[597], who published
figures and descriptions of two species, _Calamodendron striatum_ and
_C. bistriatum_.
Some years later Göppert[598], who was one of the greatest of the
older palaeobotanists, instituted another genus, _Arthropitys_[599],
for certain specimens of silicified stems from the Permian rocks of
Chemnitz in Saxony, which Cotta had previously placed in his genus
_Calamitea_ under the name of _Calamitea bistriata_[600]. Göppert
rightly decided that the plants so named by Cotta differed in important
histological characters from other species of _Calamitea_. The generic
name _Arthropitys_ has been widely adopted for a type of Calamitean
stem characterised by definite structural features. The great majority
of the petrified Calamite stems found in the English Coal-Measures
belong to Göppert’s _Arthropitys_.
[Sidenote: WILLIAMSON.]
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