Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The section diagrammatically sketched in fig. 80 is that of a Calamite
twig in which the wood appears to have been injured, and the wound has
been almost covered over by the formation of callus wood. The young
trees in a Palaeozoic forest might easily be injured by some of the
large amphibians, which were the highest representatives of animal
life during the Carboniferous period, just as our forest trees are
often barked by deer, rabbits, and other animals. Fissures might also
be formed by the expansion of the bark under the heating influence of
the sun’s rays[618]. Such a specimen as that of fig. 80 gives an air
of living reality to the petrified fragments of the Coal period trees.
It is well known how a wound on the branch of a forest tree becomes
gradually overgrown by the activity of the cambium giving rise to a
thick callus, which gradually closes over the wounded surface in the
form of two lips of wood which finally meet over the middle of the
scar. The two lips of callus are clearly shown in the fossil branch
arching over the tear in the wood just beyond the ring of carinal
canals. The tissue external to the wood represents the imperfectly
preserved cortex. A section which was cut parallel to that of fig. 80
shows a continuous band of wood beyond the wound, and the latter has
the form of a small triangular gap; this section appears to have passed
across the wound where it was narrower and has already been closed over
by the callus. The formation of a rather different kind of callus wood
has been described by Renault[619] and by Williamson and Scott[620],
in stems where aborted or deciduous branches have been overgrown and
sealed up by cambial activity.
[Illustration: FIG. 80. Diagrammatic sketch of a transverse section of
a Calamite twig, showing callus wood. From a specimen in the
Cambridge Botanical Laboratory Collection. × _ca._ 10.]
[Illustration: FIG. 81. _Calamites._ Longitudinal section (_R_, radial;
_T_, tangential) of a small branch. _b_, position of a lateral
branch. From a specimen (no. 1937) in the Williamson Collection.
Slightly enlarged.]
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