Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Some of the features to be noticed in longitudinal sections of Calamite
stems have already been described, at least as regards younger
branches. The specimen shown in fig. 81 illustrates the general
appearance of a stem as seen in tangential and radial section. In the
lower portion, _T_, the course of the vascular bundles is shown by
the black lines which represent the xylem tracheids, bifurcating and
usually alternating at each node. Between the xylem strands are the
broad principal medullary rays. At _b_ a branch has been cut through
on its passage out from the parent stem, just above the nodal line. In
tangential sections of Calamite stems one frequently sees both branches
and leaf-trace bundles (fig. 83, _A_), passing horizontally through
the wood and enclosed by strongly curved and twisted tracheids. In the
upper part of the figure (81, _R_), the section has passed through the
centre of the stem, and the wood is seen in radial view; each node is
bridged across by a diaphragm of parenchymatous cells capable of giving
rise to a surface layer of periderm[621].
An outgoing branch, as seen in a tangential section of a stem, consists
of a parenchymatous pith surrounded by a ring of vascular bundles, in
which the characteristic carinal canals have not yet been formed, but
if the section has cut the branch further from its base, there may be
seen a circle of irregular gaps marking the position of the carinal
canals. Such gaps are often occupied by thin parenchyma, and contain
protoxylem elements. The outgoing branches, as seen in a tangential
section of a Calamite stem, are seen to be connected with the wood of
the parent stem by curved and sinuous tracheids, which give to the
stem-wood a curiously characteristic appearance[622], as if the xylem
elements had been pushed aside and contorted by the pressure of the
outgoing member. A tangential section through a Pine stem[623] in the
region of a lateral branch presents precisely the same features as in
_Calamites_. The branches are given off from the stem immediately above
a node and usually between two outgoing leaf-trace bundles.
[Sidenote: RHIZOME OF CALAMITES.]
Public-domain text, read in full here on John Shaqi.
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