Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
It is an almost hopeless task to attempt to connect the various forms
of foliage-shoots with their respective stems, and to determine what
particular anatomical features characterised the plants bearing
these various forms of shoots. We may adopt Schimper’s generic name
_Calamocladus_ in the same sense as _Asterophyllites_, but as including
such other foliage-shoots as we have reason to believe belonged to
_Calamites_. Those leaf-bearing branches which conform to the type
known as _Annularia_ are however not included in _Calamocladus_, as
we cannot definitely assert that these foliage-shoots belong in all
cases to Calamitean stems. Grand’Eury’s use of _Calamocladus_ in a
more restricted sense is inadvisable as leading to confusion, seeing
that this name was originally defined in a more comprehensive manner
as including Calamitean leaf-bearing branches generally. We may define
_Calamocladus_ as follows:—
Branched or simple articulated branches bearing whorls of uni-nerved
linear leaves at the nodes; the leaves may be either free to the
base or fused basally into a cup-like sheath (_e.g._ Grand’Eury’s
_Calamocladus_). The several acicular linear leaves or segments which
are given off from the nodes spread out radially in an open manner in
all directions; they may be either almost at right angles to the axis
or inclined at different angles. Each segment is traversed by a single
vein and terminates in an acuminate apex.
As a typical example of a Calamitean foliage-shoot the species
_Calamocladus equisetiformis_ (Schloth.) may be briefly described. The
synonymy of the commoner species of fossil plants is a constant source
of confusion and difficulty; in order to illustrate the necessity of
careful comparison of specimens and published illustrations, it may be
helpful to quote a few synonyms of the species more particularly dealt
with. The exhaustive lists drawn up by Kidston in his _Catalogue of
Palaeozoic plants in the British Museum_ will be found extremely useful
by those concerned with a systematic study of the older plants.
[Illustration: FIG. 87. _Calamocladus equisetiformis_ (Schloth.).
From a specimen in the British Museum (McMurtrie Collection, no. V.
2963). _ca._ ⅓ nat. size.]
_Calamocladus equisetiformis_ (Schloth.). Fig. 87.
1809. _Phytolithus_, Martin[648].
1820. _Casuarinites equisetiformis_, Schlotheim[649].
1825. _Bornia equisetiformis_, Sternberg[650].
1828. _Asterophyllites equisetiformis_, Brongniart[651].
1836. _Hippurites longifolia_, Lindley and Hutton[652].
1855. _Calamites equisetiformis_, Ettingshausen[653].
1869. _Calamocladus equisetiformis_, Schimper[654].
1869. _Annularia calamitoides_, Schimper[654].
The above synonyms do not exhaust the list[655], but they suffice to
illustrate the necessity of a careful comparison in drawing up tables
of species, in connection with geographical distribution or for other
purposes.
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