Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In attempting to identify pith-casts the student must keep in view
the probable differences presented by the branching rhizome, the main
aerial branches and the finer shoots of the same individual. The long
internodal ridges of some casts may be mistaken for the parallel veins
of such a leaf as _Cordaites_, a Palaeozoic Gymnosperm, if there are
no nodes visible on the specimen. The fossil figured by Lindley and
Hutton[755] as _Poacites_, and regarded by them as a Monocotyledon,
is no doubt a portion of a Calamite with very long internodes. An
interesting example of incorrect determination has recently been
pointed out by Nathorst[756] in the case of certain casts from Bear
Island, originally described by Heer as examples of _Calamites_; the
vertical rows of leaf-trace casts on a _Knorria_ were mistaken for the
ribs of a Calamite stem. The specimens in the Stockholm Museum fully
bear out Nathorst’s interpretation. The undulating course of internodal
ridges and grooves is not in itself a character of specific value. If
a Calamite stem were bent slightly, the wood and medullary-ray tissues
on the concave side might adapt themselves to the shortening of the
stem by becoming more or less folded, and a cast of such a stem would
show undulating ridges and grooves on one side and straight ones on the
other[757].
A convenient classification of Calamite casts was proposed by Weiss
in 1884, founded chiefly on the number and manner of occurrence of
branch-scars—or rather branch-depressions—on the surface of pith-casts.
Weiss[758] recognised the imperfection of his proposed grouping,
and Zeiller[759] has also expressed reasonable doubts as to the
scientific value of such group-characters. Weiss instituted three
subgenera—_Calamitina_, _Eucalamites_ and _Stylocalamites_, which are
made use of as convenient terms in descriptive treatment of Calamite
casts. The following account of a few of the more typical casts may
serve to illustrate the methods employed in the description of such
specimens; the synonomy given for the different species is not intended
to be complete, but it is added with a view to drawing attention to the
necessity for careful comparison in systematic work.
A. _Calamitina._
[Illustration: FIG. 99. _Calamites_ (_Calamitina_) _Göpperti_ (Ett.).
_b_, branch scars.
From a specimen in the Manchester Museum, Owens College. ¼ nat.
size.]
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