Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This sub-genus of _Calamites_, as instituted by Weiss[760], includes
Calamitean stems or branches, which are characterised by the periodic
occurrence of branch-whorls usually represented by fairly large oval
or circular scars just above a nodal line (figs. 99, 100 and 101).
The branch-scars may form a row of contiguous discs, or a whorl may
consist of a smaller number of branches which are not in contact
basally. A form described by Weiss as _C. pauciramis_, Weiss[761],
has only one branch in each whorl, as represented by a single large
oval scar on some of the nodes of the cast. A stem of this form is by
no means a typical _Calamitina_, but it serves to show the existence
of forms connecting Weiss’ sub-genera _Calamitina_ and _Eucalamites_.
The number of internodes and nodes between the branch whorls varies
in different specimens, and is indeed not constant in the same
plant. Each nodal line bears numerous elliptical scars which mark
the points of attachment of leaves; each branch-whorl is situated
immediately above a node, and in some forms this nodal line pursues
a somewhat irregular course across the stem, following the outlines
of the several branch-scars[762]. The surface of the internodes is
either perfectly smooth or it is more frequently traversed by short
longitudinal ridges or grooves probably representing fissures in the
bark of the living stem; these are indicated by lines in fig. 99 and
by elongated elliptical ridges in fig. 101. On young stems the leaves
are occasionally found in place, as for example in an example figured
by Weiss[763] (_C. Göpperti_), or we may have leaf-verticils still in
place in much older and thicker branches[764] (cf. fig. 85, p. 330).
It occasionally happens that the bark of _Calamitina_ stems has been
preserved as a detached shell[765] reminding one of the sheets of Birch
bark often met with in forests, the separation being no doubt due in
the fossil as in the recent trees to the manner of occurrence of the
cork-cambium.
In a few cases branches have been preserved still attached to a stem
or branch of higher order; examples of such specimens are figured
by Lindley and Hutton[766], Stur[767], and others. Grand’Eury[768]
has given an idealised drawing of a typical _Calamitina_ bearing a
whorl of branches with the foliage and habit of _Asterophyllites
equisetiformis_. The specimen on which this drawing is based is in the
Natural History Museum, Paris; it shows Asterophyllitean branches in
organic connection with a Calamitean stem, but it is not quite clear if
the stem is a true _Calamitina_. A large drawing of this interesting
specimen is given by Stur[769] in his monograph on _Calamites_, also
a smaller sketch by Renault[770] in his _Cours de botanique fossile_.
Similar branches of the _Asterophyllites_ type attached to an undoubted
_Calamitina_ are figured also by Lindley and Hutton. There is, in
short, good evidence that stems of this sub-genus bore branches with
Asterophyllitean shoots.
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