Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The wood of stems of the Calamitina group of _Calamites_, in some
instances at least, was of the _Arthropitys_ type; this has been
shown to be the case in some French specimens from the Commentry
coal-field[771] and in others described by Stur[772]. The pith-casts
of _Calamitina_ are characterised by comparatively short internodes
separated by deep nodal constrictions, as shown in fig. 100. From
Permian specimens from Neu Paka in Bohemia, described by Stur[773], we
learn that there were the usual Calamite diaphragms bridging across
the wide pith-cavity at each node. Such a cast as that shown in fig.
100 is often referred to as _Calamites approximatus_ Brongn.; the
length of the internodes and the periodic occurrence of branch-scars
in the form of circular or oval depressions along a nodal line
enable us to recognise the _Calamitina_ casts. Weiss[774] points out
that in pith-casts of this form the branch-scars occur on the nodal
constriction, and not immediately above the node as is the case on
the surface of a typical _Calamitina_. This distinction is however
of little or no value; the point of attachment of a branch may be
above the nodal line, while on the pith-cast of the same stem the
point of origin of the vascular bundles of the branch is on the nodal
constriction[775].
The specimen shown in fig. 100 illustrates the appearance of a
_Calamitina_ cast. There is a verticil of branch-scars on the lowest
nodal constriction; on the right of the pith-cast the broad band of
wood is faintly indicated by the smooth surface of the rock (_x_).
Other examples demonstrating the existence of a broad woody cylinder in
_Calamitina_ stems have been figured by Weiss[776] and other writers,
and some good examples may be seen in the British Museum.
[Illustration: FIG. 100. _Calamites_ (_Calamitina_) _approximatus_
Brongn. Lower Coal-Measures of Ayrshire.
_x_, impression of the wood.
(From a specimen in the collection of Mr R. Kidston.)]
We have so far noticed the connection of certain forms of pith-casts
(_e.g._ _Calamites approximatus_), and Asterophyllitean shoots with
stems of the sub-genus _Calamitina_.
As regards the strobili our knowledge is far from satisfactory.
Stur[777] figures some fertile branches bearing long and narrow
strobili, either _Palaeostachya_ or _Calamostachys_, in close
association with _Calamitina_ stems, and Renault and Zeiller[778] give
illustrations of the association of _Calamitina_ stems with large
strobili of the _Macrostachya_ form.
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