Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Since the microscopical examination of fossil plant-anatomy was
rendered possible, a more correct interpretation of decorticated and
incomplete specimens has been considerably facilitated. The examination
of tangential sections taken at different levels in the cortex of such
a plant as _Lepidodendron_ brings out the distribution of thin and
thick-walled tissue. Regularly placed prominences on such a stem as the
_Knorria_ shown in fig. 23 are due to the existence in the original
stem of spirally disposed areas of thin-walled and less resistant
tissue; as decay proceeded, the thinner cells would be the first to
disappear, and depressions would thus be formed in the surrounding
thicker walled and stronger tissue. If the stem became embedded in mud
or sand before the more resistant tissue had time to decay, but after
the removal of the thin-walled cells, the surrounding sediment would
fill up the depressions and finally, after the complete decay of the
stem, the impression on the mould or on the cast, formed by the filling
up of the space left by the stem, would have the form of regularly
disposed projections marking the position of the more delicate tissues.
The specimen represented in the figure is an exceedingly interesting
and well preserved example of a Coal-Measure stem combining in itself
representatives of what were formerly spoken of as distinct genera.
[Illustration: FIG. 23. A dichotomously branched Lepidodendroid stem
(_Knorria mirabilis_ Ren. and Zeill.). After Renault and
Zeiller[147]. (¼ nat. size.) The original specimen is in the
Natural History Museum, Paris.]
The surface of the fossil as seen at _e_ affords a typical example of
the Knorria type of stem; the spirally disposed peg-like projections
are the casts of cavities formed by the decay of the delicate cells
surrounding each leaf-trace bundle on its way through the cortex of
the stem. The surface _g_ exhibits a somewhat different appearance,
owing to the fact that we have the cast of the stem taken at a slightly
different level. The surface of the thick layer of coal at a shows
very clearly the outlines of the leaf-cushions; on the somewhat deeper
surfaces _b_, _c_ and _d_ the leaf-cushions are but faintly indicated,
and the long narrow lines on the coal at _c_ represent the leaf-traces
in the immediate neighbourhood of the leaf-cushions.
[Sidenote: IMPERFECT CASTS.]
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