Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The first account of the anatomy of _Halonia_ we owe to Dawes[312];
this was followed by a fuller description by Binney[313]. The
history of our knowledge of this type of branch has been given
by Carruthers[314], who expressed the opinion that Halonia is
merely a fertile condition of _Lepidophloios_ and possibly of other
lepidodendroid plants. He was also inclined to regard the Halonial
tubercles as younger stages of the larger scars characteristic of the
genus _Ulodendron_. Williamson’s contributions to our knowledge of
_Halonia_ are of primary importance; he supplied further proof of the
Lepidodendroid nature of these branches and advanced our knowledge of
their anatomy. In an early paper[315] he expressed the view that the
differences on which _Halonia_ and _Ulodendron_ are separated are such
as result from a difference in age and are not of generic importance.
In the last memoir, of which he was sole author, published by the Royal
Society[316], Williamson brought forward further evidence in support of
this well-founded opinion.
That the fossils known as _Halonia_ are branches of a lepidodendroid
plant is at least certain, and it is probable that the lateral
branches which they bore were fertile, though satisfactory proof of
this is lacking. We know also that Halonia branches are characterised
by the Lepidophloios form of leaf-cushion; there is, however, no
sufficient reason to assume that such branches were never attached
to stems with the cushions of the Lepidodendron form. The further
question, namely whether Williamson was correct in his contention as
to the absence of any essential distinction between _Ulodendron_ and
_Halonia_, does not admit of an unchallenged answer. In 1903 Weiss[317]
described the anatomy of a specimen of a biseriate _Halonia_ branch of
_Lepidophloios_. The form of the leaf-cushions is unfortunately not
very well preserved, but Weiss figures other specimens with two rows
of tubercles on which the leaf-cushions are sufficiently distinct to
justify a comparison with those of _Lepidophloios_. He believes with
Williamson that it is the presence of tubercles in place of scars which
distinguishes _Halonia_ from _Ulodendron_, and that the arrangement of
the tubercles or scars is a matter of little importance. He expresses
the opinion justified by the evidence available that the absence or
presence of tubercles is merely due to accidents of preservation or,
one may add, to difference in age. Kidston[318] dissents from Weiss’s
description of his specimen as a biseriate _Halonia_; he regards it as
a Ulodendron branch of _Sigillaria discophora_ (König). Until specimens
with more clearly preserved external features are forthcoming it is
impossible to settle the point in dispute, but on the facts before
us there would seem to be a _prima facie_ case in favour of Weiss’s
contention.
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