Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In 1883 Williamson[308] described a specimen, now in the Leeds
Museum, which convinced him that _Halonia_ is merely a special form
of _Lepidodendron_ concerned with the production of fertile shoots or
strobili. Feistmantel[309] also recognised that _Halonia_ _regularis_
is identical in the form of the cushions with the type known as
_Lepidophloios laricinus_. It is worthy of note that under the name
_Halonia_, Feistmantel[310] figured a piece of decorticated axis
characterised by two rows instead of the usual spiral series of large
cup-shaped scars. Recent researches have, however, tended to break down
the distinction between _Ulodendron_ and _Halonia_ founded respectively
on the biseriate and spiral arrangement of the scars or tubercles.
The interpretation of Halonial branches as cone-bearing members of
Lepidodendroid plants has passed into a generally accepted statement
of fact, but, so far as I know, only one specimen has been figured in
which strobili are seen attached to an Halonia axis. This specimen,
described by Grand’Eury[311] from the coal-field of Gard, is hardly
sufficiently well-preserved to constitute a demonstration of the
correctness of the generally received view, which, as is not unusual,
has been repeated by one writer after another without due regard being
paid to the nature of the evidence on which the statement is based. It
may, indeed, be correct to describe Halonial branches as cone-bearing,
but there are certain considerations which make one pause before
unhesitatingly accepting this explanation. The vascular strand which
passes from the central cylinder of the shoot to the tubercle or scar
is composed of a solid rod of xylem distinguished from the main stele
by the absence of a pith. In such petrified peduncles as have been
discovered the stele is of the medullated type. The common occurrence
of strobili terminating slender branches of lepidodendroid plants,
though not a fatal objection to their attachment to Halonial shoots,
shows that in many cases the cones were borne at the tip of leafy
shoots. It may be that some of the Halonial scars are in origin like
those of the Ulodendron axes of _Bothrodendron_ and mark the position
of deciduous vegetative branches.
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