Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
_Lepidodendron Veltheimianum_ is represented by casts of stems,
the largest of which hitherto described reaches a length of 5·22
metres with a maximum diameter of 63 cm.; this specimen, figured
by Stur[371], consists of a tapered main axis giving off smaller
lateral shoots, some of which exhibit dichotomous branching. Fig.
185, C and D, represent the external features of a well-preserved
cast and impression respectively. Oblique rows of prominent cushions
wind round the surface of the stem and branches: each cushion is
prolonged upwards and downwards in the form of a narrow ridge with
sloping sides which connects adjacent cushions by an ogee curve. At
the upper limit of the broader kite-shaped portion of the cushion
the ligular pit forms a conspicuous feature; immediately below this
is the leaf-scar with its three small scars,—the lateral parichnos
strands and the central leaf-trace. The two oval areas shown in fig.
185, D, just below the lower edge of the leaf-scars, represent the
parichnos arms which impinge on the surface of the cushions on their
way to the leaves, as explained on a previous page. It is possible
that these areas were visible on the living stem as strands of loose
parenchyma comparable with the lenticel-like pits on the stipules of
_Angiopteris_[372] and the leaf-bases of Cyatheaceous ferns, or it may
be that their prominence in the specimen before us is the result of
the decay of a thin layer of superficial cortex which hid them on the
living tree. Fig. 185, B, illustrates the appearance of a stem in a
partially decorticated condition (_Bergeria_ state). A further degree
of decortication is seen in fig. 185, A, which represents the _Knorria_
condition.
[Illustration: FIG. 185. _Lepidodendron Veltheimianum._ From specimens
in Dr Kidston’s Collection. (Approximately nat. size.)]
Fig. 157 shows a Ulodendron axis of this species; in the lower part the
specimen illustrates the partial obliteration of the surface features
as the result of the splitting of the outer bark consequent on growth
in thickness of the tree. By an extension of the cracks, shown in an
early stage in fig. 157, the leaf-cushions would be entirely destroyed
and the surface of the bark would be characterised by longitudinal
fissures simulating the vertical grooves and ridges of a Sigillarian
stem. The large stumps of trees shown in the frontispiece to Volume I.
are probably, as Kidston[373] suggests, trunks of _L. Veltheimianum_ in
which the leaf-cushions have been replaced by irregular longitudinal
fissures. In old stems of _Sigillaria_ the enlarged parichnos areas
constitute a characteristic feature (p. 205), but it does not follow
that the absence of large parichnos scars is a distinguishing feature
of all _Lepidodendra_.
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