Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Under this specific name are included strobili from Upper Carboniferous
rocks which, in spite of minor differences, may be considered as one
type. The cylindrical cones vary considerably in size, some reaching a
length of 50 cm. or more. The sporophylls are attached by a pedicel,
4–8 mm. long, at right angles to the axis, while the distal portion
forms an oval lanceolate limb 10–20 mm. in length. The sporangia are
4–8 mm. long.
The branched example figured by Lindley and Hutton[419] as a variety
(L. _ornatus_ var. _didymus_) illustrates a phenomenon not uncommon in
both Palaeozoic and recent lycopodiaceous strobili.
[Illustration: FIG. 191. _Lepidostrobus._
A–D. _L. oldhamius._
B, C, D. From sections in the Binney Collection, Cambridge.
E. Megaspore. (After Kidston.)
F. Megaspore (Coal-Measures, Halifax). (After Williamson.)
G. Megaspore of _Lepidostrobus foliaceus_. (After Mrs Scott.)
H. Tangential section of sporangium. (After Bower.)
I. Part of sporangium wall, _Sm_, of the cone of _Lepidodendron
Veltheimianum_, enclosing two megaspores. (Cambridge Botany
School.)]
ii. _Lepidostrobus oldhamius_ Williamson[420]. Fig. 191, A–D.
Williamson[421] instituted this term for strobili previously
described by Binney[422], without adequate evidence, as the cones of
_Lepidodendron Harcourtii_. In shape and in the main morphological
features this type resembles _L. variabilis_, which is however known
only in the form of casts and impressions. A cone of _L. oldhamius_,
2–3 cm. in diameter, possesses a medullated stele consisting of a
ring of primary xylem (fig. 191, D, _x_) with exarch protoxylem and
no secondary elements. Maslen found several short tracheae at the
periphery of the xylem and states that these led him to compare the
cone with the vegetative shoots of _Lepidodendron vasculare_, but
the common occurrence of such elements in different types of shoot
renders them of little or no specific value. The inner cortex is like
that of vegetative shoots of _Lepidodendron_ and the middle cortex,
which was no doubt of the type described in _Lepidostrobus Brownii_,
is represented by a gap in the sections, beyond which is the stronger
outer cortex (fig. 191, D) passing into the horizontal pedicels of the
sporophylls. The section of the axis reproduced in fig. 191, D, was
figured by Binney[423] as _Lepidodendron vasculare_. The leaf-traces,
several of which are seen in the middle cortical region in fig. D,
_lt_, consist of a strand of scalariform tracheae, with a mesarch
protoxylem, succeeded by a few parenchymatous cells; beyond these there
is usually a small gap which was originally occupied by a strand of
thin-walled cells. It is important to note that in one sporophyll-trace
figured by Maslen[424] there is a strand of thin-walled elongated
elements abutting on the xylem, which he describes as phloem. This
tissue is certainly more like true phloem than any which has hitherto
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