been described in the leaf-traces of vegetative shoots. The state of preservation is not, however, sufficiently good to enable us to recognise undoubted phloem features. In such cones as I have examined no tissue has been seen which shows the histological features characteristic of the secretory zone of vegetative shoots: the “phloem” (Maslen) occupies the position in the sporophyll bundle which in the vascular bundles of foliage leaves is occupied by a dark-celled and partially disorganised tissue in continuity with the secretory zone of the main stele. It may be that in the strobili this tissue occurred in a modified form, but even assuming that the section figured by Maslen shows true phloem, an assumption based on slender evidence, this is not sufficient justification for the application of the term phloem to a tissue occupying a corresponding position in vegetative shoots and distinguished by well-marked histological features. The sporophyll-traces, as seen in the outer cortex in fig. 191, D, are partially surrounded by a large crescentic space, _p_, which was originally occupied by the parichnos. The sporangia are attached along the middle line of the sporophyll and, as in _Lepidostrobus Brownii_, a cushion of parenchyma projects into the lower part of the sporangial cavity (fig. 191, A, _a_; C, _a_). The diagrammatic sketch of part of a section in the Binney Collection reproduced in fig. 191, B, shows the position of the ligule, _l_. No megaspores have been discovered in any specimens of this type; the microspores, which occur both singly and in tetrads, have a length of 0·02–0·03 mm. The drawing shown in fig. 191, A, based on a section in the Binney Collection, illustrates the general arrangement of the parts of a typical _Lepidostrobus_. I have made use of this sketch instead of that given by Maslen, as his figure conveys the idea that the sporophylls are superposed, whereas, whether they are verticillate or spiral, a radial longitudinal section would not cut successive sporangia in the same plane. iii. _Lepidostrobus Brownii_ (Brongn.). In 1843 a specimen of a portion of a petrified cone was purchased by the British Museum, assisted by the Marquis of Northampton and Robert Brown, for £30 from a French dealer. This fossil, from an unknown locality, was briefly described by Brown in 1851[425] and named by him _Triplosporites_, but in a note added to his paper he expressed the opinion that the generic designation _Lepidostrobus_ would be more appropriate. Brongniart afterwards named the cone _Triplosporites_ _Brownii_[426], and Schimper[427] described it in his _Traité_ as _Lepidostrobus Brownii_. The type-specimen is preserved in the British Museum and the Paris Museum possesses a piece of the same fossil.