Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In the _Antediluvian Phytology_, Artis[448] figured a fossil from the
Carboniferous sandstones of Yorkshire which he called _Euphorbites
vulgaris_ on account of a superficial resemblance to the stems of
existing succulent Euphorbias. Rhode[449] also compared Sigillarian
stems with those of recent _Cacti_. The specimen described by Artis
is characterised by regular vertical and slightly convex ribs bearing
rows of leaf-scars in spiral series, like those on the cushions of
_Lepidodendron_. A few years earlier Brongniart[450] had instituted the
genus _Sigillaria_[451] for plants with ribbed but not jointed stems
bearing “disc-like impressions” (leaf-scars) disposed in quincunx; the
type-species named by the author of the genus _Sigillaria scutellata_
is identical, as Kidston[452] points out, with _Euphorbites vulgaris_
of Artis and with the plant afterwards figured by Brongniart as _S.
pachyderma_[453]. Brongniart in 1822 figured another type of stem
characterised by the absence of ribs and by prominent spirally arranged
cushions bearing relatively large leaf-scars like the upper part of
the specimen shown in fig. 203; this he named _Clathraria Brardii_, a
well-known and widely distributed Carboniferous and Permian species
now spoken of as _Sigillaria Brardi_ (figs. 196, A–C; 203). A third
type of stem figured by Brongniart as _Syringodendron striatum_[454]
agrees with _Sigillaria scutellata_ in having ribs, but differs in the
substitution of narrow oval ridges or depressions for leaf-scars; this
is now recognised as a partially decorticated _Sigillaria_, in which
the vascular bundle of each leaf is represented by a narrow ridge or
depression. The name _Syringodendron_, originally used by Sternberg, is
conveniently applied to certain forms of Sigillarian stems which have
lost their superficial tissues. A fourth generic name, _Favularia_, was
instituted by Sternberg[455] for Sigillarian stems with ribs covered
with contiguous leaf-scars of hexagonal form and prominent lateral
angles (fig. 193, A; fig. 200, G).
[Illustration: FIG. 193.
A. _Sigillaria elegans_ Brongn.
B. _Sigillaria rugosa_ Brongn. Middle Coal-Measures.
C. _Omphalophloios anglicus_ Kidst. Barnsley.
D. _Sigillaria elegans_ Brongn.
E. _Sigillaria tessellata_ Brongn.
(A, B, C, E, about ¾ nat. size. Dr Kidston’s Collection.)]
The generic or sub-generic title _Rhytidolepis_, also instituted
by Sternberg, is applied to ribbed Sigillarian stems such as _S.
scutellata_, _S. rugosa_ (fig. 193, B), _S. mammillaris_ (fig. 195),
or _S. laevigata_ (fig. 196, D). Goldenberg[456] proposed the name
_Leiodermaria_ for smooth Sigillarian stems with leaf-scars not in
contact with one another (fig. 196, C).
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