Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Immediately above the centre of the upper boundary of a Sigillarian
leaf-scar a ligule pit may often be detected, as shown in fig. 195,
_l_, and in some cases, e.g. a specimen figured by Germar[463] (fig.
196, A) as _Sigillaria spinulosa_ (identical with _S. Brardi_), some
circular scars with a central pit surrounded by a raised rim occur
on the surface of the stem, either singly or in pairs, near the
leaf-scars; these, it is suggested, may represent the position of
adventitious roots or, as Germar thought, of some deciduous spinous
processes. The leaf-scars are frequently hexagonal in shape, with the
lateral angles either rounded (fig. 200, F) or sharply pointed (fig.
200, G, H); each scar bears three smaller scars as in _Lepidodendron_,
a central circular, oval or crescentic leaf-trace scar and larger oval
or slightly curved scars formed by the two parichnos arms (fig. 195,
_p_). The larger size of the parichnos arms, the individual cells of
which may often be detected as a fine punctation, is a distinguishing
feature of the genus, but otherwise the structure is very similar to
that in _Lepidodendron_. As shown in figs. 195, 200, F, G, the three
scars may occur nearer the upper than the lower margin of the leaf-base
area.
_Lepidodendron Wortheni_[464] (fig. 196, E), described from North
America by Lesquereux[465], by Zeiller[466] from France, and by
Kidston[467] from the Upper and Middle Coal-Measures of England,
may be quoted as a _Lepidodendron_ bearing a close resemblance to
_Sigillaria_. The shoots bear cushions two or three times as long
as broad and without the usual median division, but with numerous
irregular and discontinuous transverse wrinklings. _Lepidodendron
Peachii_ Kidston[468] affords another example of a form agreeing both
with _Sigillaria_ and with _Lepidodendron_. An Upper Devonian type
described by White[469] as _Archaeosigillaria primaeva_ affords a
striking instance of the combination on one stem of Sigillarian and
Lepidodendroid leaf-cushions.
The difference between the original surface of a Sigillaria stem and
that of partially decorticated specimens is seen in figs. 196, C and
D; in fig. C the bark of _Sigillaria Brardi_ shows the characteristic
wrinklings of the superficial tissue, while at a slightly lower level
the leaf-scars are replaced by the parichnos casts, a, and fine
longitudinal striations represent the elongated phelloderm cells laid
bare by the exfoliation of the surface-layers. Similarly, in the rib
of _Sigillaria laevigata_ (fig. 196, D) the parichnos arms, _p_, and
the longitudinal striations are exposed at the lower level, while the
surface is smooth and bears rows of widely separated leaf-scars.
[Illustration: FIG. 197. _Carica sp._ From the Royal Gardens, Kew.
(Much reduced.) M.S.]
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