Fossil plants, Vol. 2 : $b A text-book for students of botany and geology — John Shaqi
Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In addition to the two types of cone already mentioned,
_Sphenophyllostachys Dawsoni_ and _S. Römeri_, others have been
described by Kidston from carbonised impressions. One of these is the
fertile branch of _Sphenophyllum majus_[8]. The basal portions of the
bracts of each whorl form a narrow collar round the axis of the cone;
the free portion of each bract consists of a lamina divided into two
equal bifid lobes bearing on its upper surface one group, or possibly
two groups, of four sessile sporangia between the narrow coherent bases
of the laminae and the sinus between the terminal lobes (fig. 113, C).
Another characteristic feature is the greater length of the internodes;
this renders the cone less compact and less sharply differentiated from
the vegetative shoots than those of other species. A specimen in Dr
Kidston’s collection illustrates the peculiar character of the fertile
portion of this species; it consists of an axis bearing a succession of
lax sporophylls succeeded above and below by whorls of sterile leaves.
In this species, therefore, we cannot speak of a compact strobilus at
the end of a shoot of limited growth, but of axes in which sterile and
fertile leaves are borne alternately[9], a condition recalling the
alternation of foliage leaves and sporophylls in _Tmesipteris_ and in
_Lycopodium Selago_.
[Illustration: FIG. 113.
A. _Sphenophyllostachys Römeri._ (Solms-Laubach.)
B. _Sphenophyllum trichomatosum_ Stur.
C. _Sphenophyllum majus._ Bronn. (A–C. After Kidston.)]
Another form of cone, also from the Middle Coal Measures, is referred
by Kidston to _Sphenophyllum trichomatosum_ Stur[10] (fig. 113, B):
this is characterised by the more horizontal position of the bracts,
which “do not appear to be so much or so suddenly bent upwards in their
distal portion as in some other species of _Sphenophyllum_,” and by
sessile sporangia borne singly on the upper face of each bract.
[Illustration: FIG. 114. _Sphenophyllostachys fertilis_ (Scott).
(After Scott.) Diagram of a node in longitudinal section, showing
one sporophyll and the base of the opposite one. _v.l._ ventral
lobe of sporophyll; _v.s._ one of the segments into which it
divides; _v.s′._ stump of another segment; _d.l._ dorsal lobe;
_d.s._, _d.s′._ segments of dorsal lobe.]
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