Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In 1889 Williamson[642] described several specimens of petrified
shoots from the Coal-Measures of Halifax which he named _Lepidodendron
mundum_: these are now known to be branches of a _Bothrodendron_. The
discovery was made by Mr Lomax[643] who found specimens showing the
external characters of _Bothrodendron_ and the anatomical characters
of _Lepidodendron mundum_. In some of the smaller twigs, the stele
consists of a solid core of xylem with external protoxylem; but in
the majority of specimens the centre of the xylem is replaced by
parenchymatous tissue, either as a small axial strand or, as in the
specimen shown in fig. 215, D, a wide pith, the elements of which
are arranged in regular vertical series. A diagrammatic section of
a small axis is represented in fig. 215, A: this branch, 2 mm. in
diameter, is composed of a broad outer cortex consisting exclusively of
primary tissue the outer cells of which are smaller and have thicker
walls than the more internal elements. The leaf-traces, _lt_, are
accompanied by a strand of delicate tissue, the parichnos. The stele
is almost solid; the tissues in contact with the xylem have not been
preserved but the inner cortex is represented by a few layers of small
parenchymatous cells, _c_¹. The larger section shown in fig. 215,
D, was cut from a specimen from Dulesgate of which the smooth surface
exhibits the characteristic leaf-scars of _Bothrodendron_. The section
measures 3 cm. in its longest diameter and the stele has a breadth of
3 mm. The outer cortex has a smooth surface and is composed of rather
thick-walled cells succeeded by a zone of secondary elements. The
middle cortex has disappeared and the space is partially occupied by
Stigmarian rootlets, _s_, and crushed patches of cortical tissue. The
position of a leaf-scar is seen at _a_; this is more clearly shown in
the enlarged drawing fig. E.
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