Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The xylem is therefore centripetal in origin as in _Sphenophyllum_
and in nearly all recent and fossil members of the Lycopodiales. In
the type-specimen of _Cheirostrobus_ the vascular cylinder of the
cone consists entirely of primary xylem, but secondary xylem has been
found in a more recently discovered specimen[17]. Secondary xylem
occurs also in the peduncle of the cone. No appreciable remains of
phloem have been found. The cortex consists of slightly elongated
rather thick-walled tissue containing secretory sacs. Crowded
superposed whorls of bracts (or sporophylls), usually twelve in each
whorl, are borne on the axis and each sporophyll receives a single
vascular bundle from one of the vertical ridges of the xylem column
(fig. 117, A, _lt_). The members of each whorl are connate at the
base: from this narrow collar each sporophyll branches into an upper
or dorsal and a lower or ventral limb (fig. 117, A, _f_ and _s_).
Each limb divides palmately at a short distance from its origin into
three slender segments, which extend in a horizontal direction and
terminate in large laminar expansions (fig. 117, B, _s_) to afford
a protective covering to the surface of the cone. The upper set of
three segments, constituting sporangiophores (fig. 117, A, B, _f_) or
fertile divisions of the sporophyll, expand distally into comparatively
bulky laminae; each of these bears on its adaxial face four diagonally
placed outgrowths which form the short pedicels of very long and narrow
sporangia. The three lower segments—the sterile divisions of the
sporophylls—(fig. 117, A, B, _s_) are similar to the upper set except
in their greater length and in the kite-shaped form of their distal
laminae which are provided with lateral lobes. The single vascular
strand which supplies each sporophyll is represented at _lt_ in fig.
117, B; at _lt′_ the strand has divided into four, the three upper
bundles in the figure supply the sterile segments and the single lower
bundle ultimately divides into three which supply the fertile segments.
A pair of blunt processes (fig. A, _s_) extend downwards over the ends
of the underlying fertile lamina and two slender prolongations extend
upwards through several internodes.
[Illustration:
FIG. 117. A, B. _Cheirostrobus pettycurensis_ Scott. (After Scott.)
C, D. _Pseudobornia ursina_ Nath. (After Nathorst.)
A. Diagrammatic radial longitudinal section of part of the
cone-axis and two sporophylls. _lt_, bundle passing out to
sporophyll; _f_, fertile segment of sporophyll showing two
sporangia; _s_, sterile (lower) segment.
B. Part of transverse section. _x_, stele; _lt_, _lt′_, bundles on
their way to sporophylls; _a_, tips of sterile segments of lower
sporophylls.
C. Palmately branched leaf (½ natural size).
D. Node of stem showing leaf-bases.]
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