Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
It is not until we ascend the geological series as far as the older
Jurassic rocks that we meet with a type which can with confidence be
classed with the Schizaeaceae, as least so far as sporangial characters
are concerned. The species _Klukia exilis_ is selected as the best
known and most widely-spread representative of Jurassic Schizaeaceae.
_Klukia exilis_ (Phillips)[829]. Fig. 259.
The generic name _Klukia_ was proposed by Raciborski[830] for a species
originally described by Phillips[831] from the Inferior Oolite of
the Yorkshire coast as _Pecopteris exilis_. Bunbury’s[832] discovery
(supplemented by additional evidence obtained by Raciborski) of
well-preserved sporangia justified the substitution of a distinctive
designation for the provisional term _Pecopteris_.
[Illustration: FIG. 259. _Klukia exilis_ (Phillips). (Figs. 1–3, × 40;
fig. 4, × 3; fig. 5, nat. size.)]
The species may be defined as follows:—
Frond tripinnate, of the _Cladophlebis_ type; pinnae linear,
lanceolate, attached to the rachis at a wide angle. Ultimate
segments short and linear, entire or, in the lower part of a frond,
crenulate, 5 mm. long or occasionally longer. Sporangia 0·5 mm. in
length, borne singly on the lower surface of the lamina in a row on
each side of the midrib.
A re-examination[833] of the specimen described by Bunbury confirmed
his account of the structure of the sporangia. The pinna shown in
fig. 259 is characterised by unusually small fertile pinnules some
of which bear 10 sporangia in two rows; the annulus includes about
14 cells. Fertile specimens of this and similar forms are figured by
Raciborski[834] from Jurassic rocks of Poland, and good examples of the
English species may be seen in the Leckenby collection, Cambridge, in
the British Museum, the museums of Manchester, Scarborough, and other
places.
It is possible that specimens referred to _K. exilis_ by Yokoyama[835]
from Wealden strata in Japan may afford evidence of the persistence
of the species beyond the Jurassic era, but in view of the close
resemblance of the sterile fronds described from Wealden strata as
_Cladophlebis Brownii_[836] and _C. Dunkeri_[836] to those of _Klukia
exilis_, identity can be established only by an examination of fertile
specimens. A Jurassic fern recently described by Yabe[837] from Korea
as _Cladophlebis koraiensis_ may be identical with _K. exilis_ and
there is little doubt as to the existence of the species in Jurassic
Caucasian strata[838].
[Illustration: FIG. 260. _Ruffordia Goepperti_. (A, C, sterile; B,
fertile; slightly reduced. Specimens from the Wealden of Sussex;
British Museum; V. 2333, V. 2160, V. 2166.)]
_Ruffordia Goepperti_ (Dunk.). Fig. 260.
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