Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This species, which is characteristic of Jurassic rocks, is less
completely known than the two types described above, but in the form
and venation of the pinnae there is little difference between the
Rhaetic and Jurassic plants. The leaves of the Jurassic species appear
to have been smaller and more like those of _Dipteris conjugata_ (fig.
231); there are no indications of the existence of the two curved
arms at the summit of the petiole which form so striking a feature
in _D. exile_ and _D. Nathorsti_. No sporangia have been found on
English specimens, but it is safe to assume their agreement with those
of other species. A more complete list of records of _D. rugosum_ is
given in the first volume of the British Museum Catalogue of Jurassic
plants[950].
[Illustration: FIG. 283. _Dictyophyllum rugosum_ (Lind. and Hutt.).
(Brit. Mus. Nat. size.)]
Nathorst[951] has recently drawn attention to certain differences
between _Dictyophyllum_ and _Dipteris_. The pinnate division of the
pinnae is not represented in the fronds of the recent species, but this
method of lobing, which is a marked characteristic of _Dictyophyllum_,
is less prominent in _Clathropteris_; and in _Camptopteris lunzensis_
Stur[952], an Austrian Upper Triassic species, the pinnae are entire.
In _Dictyophyllum_ the sori cover the whole lower surface of the leaf;
in _Dipteris_ they are more widely separated and the sporangia have a
diameter of 0·02 mm., but in _Dictyophyllum_ the diameter is 0·4–0·6
mm. Moreover in _Dictyophyllum_ the sori contain 5 to 8 sporangia,
whereas in _Dipteris_ they are much more numerous. Despite these
differences it is clear, as Nathorst says, that _Dictyophyllum_,
_Clathropteris_, and _Camptopteris_ are existing types very closely
allied to _Dipteris_. It is a matter of secondary importance whether
we include all in the Dipteridinae or follow Nathorst’s suggestion and
refer the fossil genera to the separate family Camptopteridinae.
_Thaumatopteris._
This genus, founded by Goeppert[953] for a Rhaetic plant from Bayreuth,
is by some authors[954] regarded as identical with _Dictyophyllum_,
but it has recently been resuscitated by Nathorst[955] for specimens
which he names _T. Schenki_, formerly included by Schenk in his species
_T. Brauniana_[956]. It bears a close resemblance, in the long linear
pinnules with an entire or crenulate margin, to _Dictyophyllum Fuchsi_
described by Zeiller[957] from Tonkin, and it would seem hardly
necessary to adopt a distinctive generic designation. The sporangia
have a vertical or slightly oblique annulus and the rhizome is similar
to that of _Dictyophyllum exile_. The habit of the genus is shown in
fig. 284, which represents one of the German Rhaetic species.
[Illustration: FIG. 284. _Thaumatopteris Münsteri._ (From a specimen in
the Bergakademie, Berlin; ⅓ nat. size.)]
_Clathropteris._
_Clathropteris meniscoides_, Brongn. Fig. 285.
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