Other species of the genus are recorded from Liassic rocks of Steierdorf[978] (Hungary) and of Bornholm[979]. Nathorst[980] has described a small Rhaetic species from Scania: a French Permian plant described by Zeiller[981] and compared by him with _H. dichotoma_, may be a Palaeozoic example of this Dipteris-like genus. Some segments of leaves from the Eocene beds (Middle Bagshot) of Bournemouth, and now in the British Museum, described by Gardner and Ettingshausen[982] as _Podoloma polypodioides_, bear a close resemblance in the venation to the lamina of _Dipteris conjugata_. CHAPTER XXII. Marattiales (Fossil). The discovery of Pteridosperms has necessarily led to a considerable modification of the views formerly held that existing genera of Marattiaceae represent survivors of a group which occupied a dominant position in the forests of the Coal age. Mr Arber writes:—“The evidence, formerly regarded as beyond suspicion, that the eusporangiate ferns formed a dominant feature of the vegetation of the Palaeozoic period, has been undermined, more especially by the remarkable discovery of the male organs of _Lyginodendron_ by Mr Kidston. At best we can only now regard them as a subsidiary group in that epoch in the past history of the vegetable kingdom[983].” Dr Scott expresses himself in terms slightly more favourable to the view that the Marattiaceae represent the aristocracy among the Filicales. He says:—“We now have to seek laboriously for evidence, which formerly seemed to lie open to us on all hands. I believe, however, that such careful investigation will result in the resuscitation of the Palaeozoic ferns as a considerable, though not as a dominant group[984].” Zeiller’s faith[985] in the prospect of Marattiaceous ferns retaining their position as prominent members of Palaeozoic floras, though shaken, is not extinguished: he recognises that they played a subordinate part.