Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The latter author in 1891[61], in ignorance of Marion’s proposal to
adopt the name _Gomphostrobus_, instituted a genus _Psilotiphyllum_
for the sporophylls of a species originally described by Geinitz[62]
as _Sigillariostrobus bifidus_, but he subsequently adopted Marion’s
designation and with some hesitation included the French and German
specimens in the Psilotales. As stated elsewhere[63], Potonié’s
arguments in favour of his view hardly carry conviction, and it is
probably more in accordance with truth to deal with _Gomphostrobus_ in
the chapter devoted to the Coniferales.
_Psilophyton._
The generic title _Psilophyton_, instituted by the late Sir
William Dawson[64], has become familiar to geologists as that of
a Pre-Carboniferous plant characteristic of Devonian and Silurian
rocks in Canada, the United States of America, and Europe. From the
botanist’s point of view the name stands for miscellaneous remains
of plants of different types and in many cases unworthy of record.
The genus was founded on impressions of branched axes from the
Devonian strata of New Brunswick resembling the rachis and portions of
lateral pinnae of ferns or the forked slender twigs of a Lycopod. The
type-species _Psilophyton princeps_ Daws. as represented on somewhat
slender evidence in Dawson’s restoration, which accompanies the
original description of the genus and has since been copied by several
authors, is characterised by the possession of a horizontal rhizome
bearing numerous rootlets and giving off dichotomously branched aerial
shoots with spinous appendages, compared with rudimentary leaves, and
terminating in slender branchlets bearing pendulous oval “spore-cases”
from their tips. Some of the branchlets exhibit a fern-like vernation.
The plant is spoken of by Dawson as apparently a generalised type[65],
resembling in habit and in its rudimentary leaves the recent genus
_Psilotum_ and presenting points of contact with ferns. Specimens were
found in an imperfectly petrified state showing a central cylinder of
scalariform tracheae surrounded by a broad cortical zone of parenchyma
and fibrous tissue.
Among other species described by the author of the genus we need only
mention _Psilophyton robustius_, characterised by vegetative shoots
and “spore-cases” similar to those of the type-species; but, as
Solms-Laubach[66] has pointed out, the petrified sections referred by
Dawson to _P. robustius_ are of an entirely different anatomical type
from that of _P. princeps_[67].
British fossils from the Old Red Sandstone from the north of Scotland,
Orkney and Caithness, originally figured by Hugh Miller and compared
by him with algae but more especially with recent Lycopods, were
subsequently placed by Carruthers[68] in the genus _Psilophyton_ as _P.
Dechianum_, the specific designation being chosen on the ground that
the Scotch specimens are specifically identical with fossils described
by Goeppert[69] as _Haliserites Dechianus_.
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