Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Palaeobotanical literature contains numerous descriptions of Palaeozoic
petrified wood occasionally enclosing an _Artisia_ pith-cast described
under such names as _Dadoxylon_, _Cordaioxylon_, _Araucarioxylon_,
etc., and regarded as portions of Cordaitean stems. It is, however,
certain that much of this material belonged to stems other than those
of _Cordaites_. Recent research has demonstrated the insufficiency of
the secondary xylem alone, however well preserved, as a safe guide to
generic position: stems identical in the structure of the secondary
xylem differ in that of the primary portion of the stele, and it is on
the characters of the latter tissues that several genera have recently
been founded. _Mesoxylon_ affords a striking example of the importance
of the primary xylem as a distinctive feature. As Gothan[646] points
out, the species of _Calamopitys_ recently made the type of a new
genus _Eristophyton_[647] would, in the absence of the primary xylem,
probably be regarded as Cordaitean. It is important to recognise the
limitations imposed by the imperfection of the material; we cannot
in most cases determine whether a specimen should be referred to
_Cordaites_ or _Mesoxylon_, and while it may be described as probably
Cordaitean in affinity there remains the possibility that some of the
Palaeozoic plants with secondary wood like that of _Cordaites_, if
their reproductive organs were known, would not be included in the
Cordaitales. Goeppert’s species _Araucarites Tchihatcheffi_, which
Renault[648] quotes as _Cordaites_, has recently been assigned to a
new genus _Mesopitys_[649] because of certain distinctive features of
the primary xylem. Additional examples might be quoted pointing to the
tendency of recent and more thorough investigation to establish the
fact that the occurrence of Permo-Carboniferous wood of the Araucarian
type does not necessarily denote the existence of _Cordaites_. The
question of nomenclature is necessarily raised in this connexion.
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