Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In recent years it has been customary to assign Palaeozoic wood with
Araucarian pitting to the genus _Dadoxylon_, while wood of the same
general type from more recent strata is by many authors referred to
_Araucarioxylon_[650]. This arbitrary distinction based on a difference
in age is open to serious objection. Fossil wood of the Araucarian type
is widely scattered in strata ranging from Carboniferous to Jurassic
periods; it also occurs in later formations. The fact that on the one
hand Araucarian plants, as recognised by cones and foliage-shoots, are
especially characteristic of Jurassic floras and occur more rarely in
Rhaetic and Triassic floras, and on the other hand that _Cordaites_ and
its allies reached their greatest development in Permo-Carboniferous
times, renders it probable that in the majority of cases a distinctive
name based on geological age would be in accordance with botanical
differences. But we have no satisfactory data as to the upper limits
of the Cordaiteae or the lower limits of the Araucarineae: in all
probability the two families overlapped and co-existed for more than
one geological period. It is, moreover, the plants from formations
where overlapping occurred that are the most critical from a botanical
standpoint. The age-distinction is therefore at best an artificial
one and may be seriously misleading. Potonié[651] and Gothan[652]
have emphasised the desirability of adopting the name _Dadoxylon_ for
all wood of the Araucarian type irrespective of age. If a particular
specimen can be correlated definitely with _Cordaites_ or some other
genus it should be so designated, but the fragmentary nature of the
records usually precludes this simple course. The most logical plan is
to use the name _Dadoxylon_ for all woods with Araucarian characters if
there is no sufficient reason for employing a less provisional term.
If the evidence clearly points to the Araucarineae the generic name
_Araucarioxylon_ should be added in parentheses after _Dadoxylon_,
but whether or not this is done, a statement as to the geological
age of the fossil will in itself be some assistance in enabling the
student to form an opinion on the balance of probability in favour
of a Cordaitean or an Araucarian affinity. The course suggested
by Gothan[653], namely to add _Cordaites_ after _Dadoxylon_ if an
_Artisia_ pith is present, is rendered inoperative now that we know
that a discoid pith occurs in more than one genus. In this chapter we
are concerned primarily with _Cordaites_ and with such stems as may
fairly be regarded as Cordaitean: examples of fossil wood from later
formations are dealt with in another place. A distinction between
_Araucarioxylon_ and _Cordaioxylon_ stems has been based by Felix on
the nature of the pith-casts; those of the _Artisia_ type he refers to
_Cordaioxylon_, while Palaeozoic stems with _Tylodendron_ pith-casts
are assigned to _Araucarioxylon_[654]. This distinction can, however,
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