Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Though it is impossible in many cases to form an estimate of the age
of individual plants, there are clear indications that some specimens
afford notable instances of longevity. Chamberlain estimates the age of
some plants of _Dioon spinulosum_ as exceeding 400 years and mentions
an example of _D. edule_ that is probably 1000 years old. An unusually
tall plant of _Encephalartos_ in the Botanic Garden of Amsterdam
is believed by Prof. de Vries to have reached the venerable age of
2000 years[36]. The restricted range and in many cases the solitary
existence of recent Cycads, with their tall stems clothed with the
persistent cork-covered stumps of thousands of fronds, deepens the
impression of antiquity derived from a study of the geological history
of this dwindling race.
_Stems._ The tall columnar stems of some species of _Cycas_, often
branched or bearing numerous ovoid buds like enlarged bulbils[37],
are characterised by the regular alternation of large and small
leaf-bases as seen in the stem of _C. circinalis_ reproduced in fig.
380. In older stems of this species the leaf-bases are exfoliated and
the stem is covered with wrinkled and fissured cork; but in _Cycas
revoluta_ the leaf-bases are even more persistent. The columnar but
relatively stout stems of _Encephalartos_ (figs. 379, 382, 386, A) and
_Ceratozamia_ are similarly encased in a covering of petiole-bases, but
in these genera the differences between foliage-leaves and bud-scales
is much less obvious and there is no zonal alternation. On the stems
of _Macrozamia_ the rhomboidal leaf-bases are more uniform in size
and there are no scale-leaves. The tall and often palm-like stems of
_Microcycas_ sometimes show transverse rings on the bark marking
the position of former terminal buds, and in older trunks these may
disappear, leaving a fissured bark[38]. In _Cycas siamensis_ the
tuberous stem is similarly covered with a rough bark (fig. 383) and the
stems of _Zamia_ are also characterised by an absence of persistent
leaf-bases (figs. 381, B; 395, I_a_, _a_). It is pertinent to remind
the palaeobotanical student of the occurrence of flowering plants
with stems closely simulating those of some Cycads. Prof. Bower[39] in
describing _Rhynchopetalum montanum_, an Abyssinian Lobeliaceous plant,
drew attention to the similarity in surface-features and to some extent
in anatomical structure to cycadean stems. The resemblances are further
emphasised in a more recently published account of the same species
under a different name, _Lobelia Rhynchopetalum_[40].
[Illustration: Fig. 382. _Encephalartos Ghellinckii._ (¹⁄₁₁ nat.
size.)]
[Illustration: Fig. 383. _Cycas siamensis._ (From the _Encyclopaedia
Britannica_.)]
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