Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
_Encephalartos._ The fronds of this genus, in _Encephalartos
Laurentianus_[45] reaching the exceptional length of 7 metres, bear
alternate pinnae exhibiting a considerable range in form and breadth.
In _E. longifolius_, _E. Altensteinii_ (fig. 386, C), _E. Lehmanni_,
etc., the pinnae are for the most part linear, reaching a length of
20 cm. and a breadth of 2 cm.: in _E. caffer_ (fig. 387, D), _E.
latifolius_, and others the pinnae are broader and shorter and often
spinous. A frond of _E. longifolius_ or _E. Altensteinii_ may bear both
entire and lobed, spinous pinnae. In _E. Frederici-Guilielmi_ (fig.
387, G) and _E. Ghellinckii_[46] (fig. 382) the pinnae are very narrow
and almost filiform, with revolute edges. The thick and leathery pinnae
of some species are attached obliquely to the edge or to the upper
sloping sides of the rachis which forms a prominent ridge between the
rows of leaflets, and characteristic oval scars are left on the fall
of the pinnae (fig. 387, D, G′). The lamina in most species contains
several veins more or less parallel to the margins and often much more
prominent on the lower than on the upper surface.
[Illustration: Fig. 386.
A. _Encephalartos Altensteinii_, apex of stem.
B. _Dioon edule_,megastrobilus. (From a photograph by Mr S. M. Wadham.)
C. _Encephalartos Altensteinii_, frond.]
[Illustration: Fig. 387. Cycadean fronds. A, _Cycas circinalis_;
B, _Macrozamia Fraseri_; C, _Macrozamia Denisoni_; D, _Encephalartos
caffer_; E, F, _Dioon edule_ from below and above; G, _Encephalartos
Frederici-Guilielmi_, G′, side-view; H, _Ceratozamia mexicana_; I,
_Cycas circinalis_, lower part of young frond.]
_Zamia._ In _Zamia angustifolia_ (fig. 385, C) and _Z. linifolia_ the
pinnae are long and very narrow: the other extreme is represented by
_Z. Wallisii_[47] (fig. 388) with broad ovate segments reaching a
length of nearly ·5 metre and attached to the rachis by a short stalk;
the veins are prominent and dichotomously branched. Other forms of
pinnae are represented by _Z. integrifolia_, _Z. floridana_, and _Z.
Loddigesii_ (figs. 389, 390, 395). The broad and short pinnae of
_Z. furfuracea_[48] bear a close resemblance, except in the absence
of an auriculate base, to those of some species of the fossil genus
_Otozamites_. The broadly linear pinnae of _Z. pseudoparasitica_ (45
cm. × 3 cm.) often show longitudinal wrinklings on drying which suggest
comparison with the corrugated lamina of the fossil species _Nilssonia
brevis_. A basal pad or callosity on the slender bases of the pinnae is
characteristic of many _Zamia_ fronds.
[Illustration: Fig. 388. Pinna of _Zamia Wallisii_. From a drawing
after A. Braun in the Kew Herbarium. (⅓ nat. size.)]
[Illustration: Fig. 389. _Zamia integrifolia_ bearing a megastrobilus
and showing foliage-leaves and scale-leaves. A, microsporophyll; B,
megasporophyll. (After Rendle, from Jacquin.)]
[Illustration: Fig. 390. Small frond of _Zamia Loddigesii_. (⅔ nat.
size.)]
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