Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The generic name _Niponophyllum_[747] was proposed for some petrified
specimens of leaves or possibly leaflets from Upper Cretaceous beds
in Japan which, though not definitely assigned to a group or family,
are considered by the authors of the genus ‘to lie [anatomically]
somewhere between _Cordaites_ and _Cycadeoidea_’ ‘with a closer
similarity to the former than to the latter if we compare the whole
_Cordaites_ leaf with our blade.’ The data on which this conclusion
is based are, however, insufficient to justify a reference of
_Niponophyllum_ to the Cordaitales or indeed to lend any substantial
support to the opinion that the Japanese specimens are anatomically
more akin to _Cordaites_ than to other plants. The type-species is
represented by two specimens of leaf-fragments about 0·4 mm. thick and
from 6 to 9 mm. broad containing from 21 to 33 vascular bundles; the
upper part of the mesophyll is composed of palisade tissue and the
stomata appear to be confined to the lower epidermis. Each bundle is
accompanied by an =I=-shaped girder, and small patches of sclerenchyma
occur next the upper epidermis between the girders; there are no
resin-canals: the vascular bundles are collateral, the xylem is said
to be almost entirely centripetal and exarch, but in the absence of
evidence afforded by longitudinal sections the details of structure
cannot be definitely determined. A comparison is made with Cycadean
leaves and with leaves of Araucarineae and Podocarpeae, also with
_Cordaites_; another type with which _Niponophyllum_ may be compared is
_Desmiophyllum Solmsi_[748].
The genus is interesting as an example of a petrified gymnospermous
type of leaf characterised by the absence of resin-ducts and
transfusion-tracheids, the possession of collateral, apparently
exarch, bundles enclosed in a double sheath; but the data supplied
are insufficient to enable us to allocate the specimens to a position
within the class.
A specimen described by Schenk[749] as _Eolirion primigenium_ from
Lower Cretaceous beds in the Carpathian mountains closely resembles in
habit a foliage-shoot of _Poa-Cordaites_; the leaves are narrow and
linear with obtuse apices and attached, apparently, in a close spiral.
Schenk assigns the plant to the Monocotyledons, but its systematic
position must be left unsettled.
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