Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The list of Mesozoic specimens resembling _Cordaites_ leaves might
be extended. Apart from some Triassic and Rhaetic examples which may
well be Cordaitalean, there are many others which, though similar in
form and venation to _Cordaites_, are in all probability more closely
related to _Agathis_ and other genera; the species _Dammarites Bayeri_
recently described by Zeiller[750] from the Upper Cretaceous of
Bulgaria is a case in point. The Araucarian character of the wood of
_Cordaites_ precludes any satisfactory discrimination between Mesozoic
Araucarian stems and those of Cordaitalean species, at least in the
case of such material as is usually available.
=Titanophyllum.= Renault.
_Titanophyllum Grand’Euryi_ Renault. The remarkable leaves on which
this genus and species are founded[751] were discovered in the
Commentry coalfield; they occur as detached specimens and cannot be
correlated with any known stem. Renault suggests that the Autun stems
referred to _Colpoxylon_ may have borne the _Titanophyllum_ leaves,
but this correlation rests only on the dimensions of the stems and
the occurrence of transversely elongated scars on the surface. The
lamina is thick and coriaceous, 70–75 cm. long and 20–25 cm. in
breadth; the veins are parallel but not branched; numerous longitudinal
striations on the upper surface indicate the presence of hypodermal
stereome-strands; stomata are abundant on the lower surface and the
more or less rectangular cells in the neighbourhood of the stomata
appear to be papillose (fig. 485, A, B). The distal region of the
lamina is often torn into strips (fig. 485, A); the approximately
rectangular leaf has a broad elliptical base 9–10 × 3–4 cm.
[Illustration: Fig. 485. _Titanophyllum Grand’Euryi._ A, leaves
¹⁄₃₀ nat. size; B, stomata on the lower surface of the lamina. (After
Renault and Zeiller.)]
Dr White[752] describes a specimen from the Lower Coal Measures of
Missouri as ? _Titanophyllum Brittsii_ which he speaks of as the thick
base of a leaf similar to that described by Renault but, as White says,
no formal diagnosis is possible without more satisfactory material.
Such evidence as is available suggests that _Titanophyllum_ is a type
of Cordaitalean leaf probably closely allied to _Cordaites_.
CHAPTER XXXIV.
CORDAITALES.
C. =PITYEAE.=
=Pitys=. Witham.
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