Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This generic name in the form _Pitus_ was first used by Witham[753]
for some petrified pieces of large stems from Lower Carboniferous
strata in the south of Scotland (Berwickshire). In an earlier paper
Witham[754] gives an account of the beds on the banks of the Tweed from
which the specimens of _Pitys_ and other plants were obtained. The name
was revived by Goeppert[755] as a substitute for _Pissadendron_[756]
adopted by Endlicher and Unger: Scott[757] employed the older name in
the account of his investigations, which have added greatly to our
knowledge of Witham’s types. The distinguishing characters of _Pitys_
are: the Araucarian pitting of the secondary xylem, the large pith, the
occurrence of numerous small primary xylem strands of mesarch structure
in the peripheral region of the pith, and the simple nature of the
leaf-traces. Nothing is known of the extrastelar tissues, and beyond
the fact that the species were arborescent we have no information with
regard to the foliar[758] or reproductive organs.
[Illustration: Fig. 486. _Pitys antiqua._ Central region of stem,
showing primary xylem strands as black dots. (Kidston Collection, 598
A.)]
_Pitys antiqua_ Witham.
In 1899 Scott[759] published a preliminary description of some sections
in the possession of Dr Kidston from Lennel Braes (Berwickshire:
Calciferous series) under the name _Araucarioxylon antiquum_, but
in the more complete account the name _Pitys_[760] is adopted. The
following description is based on Scott’s paper and on the examination
of the sections. Fig. 486 represents a diagrammatic sketch of a section
(2·7 × 2·7 cm.) of the central region of a stem which had lost most
of its secondary wood. In some cases the pith reaches a diameter of
3 cm.; it consists of large parenchymatous cells broader than deep,
several of which contain a black substance and are probably secretory
elements: the intercellular spaces are fairly large. Horizontally
extended gaps, due to shrinkage and collapse, occur in the pith and
give to it an appearance not unlike that of _Cordaites_. Numerous small
elliptical strands of primary xylem, 0·15–0·3 mm. in diameter, are
scattered in the peripheral region (fig. 486) at varying distances from
the inner edge of the secondary wood and occasionally, when about to
bend outwards as a leaf-trace, a primary xylem-strand abuts directly
on the secondary xylem. The protoxylem occupies a more or less central
position in each mesarch strand (fig. 487, B, _px_): the centrifugal
tracheids have the Araucarian pitting while the centripetal elements
are spiral. A few parenchymatous cells are associated with the more
central tracheids. A leaf-trace about to bend outwards into the zone of
secondary wood is double, but at a slightly higher level in its course
it assumes the form of a single strand. The foliar bundles are thus
single and not true double traces. Scott recognised, at the extreme
edge of the pith, an association with outgoing leaf-traces of a single
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