Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Miss Elkins and W. Wieland[774] have recently described some Upper
Devonian wood from Indiana characterised by a grouping of the circular
or elliptical bordered pits in the radial walls of the tracheids
similar to those in _Callixylon Trifilievi_ which they include
together with the Middle Devonian species _Cordaites Newberryi_[775]
in Zalessky’s genus. Though these two American species are comparable
in the discontinuous arrangement of the tracheal pits with the
Russian type the latter is characterised by the presence of primary
xylem-strands, a feature that has not been recognised in the American
stems: it would seem, then, undesirable to adopt the designation
_Callixylon_ in preference to _Dadoxylon_ unless there is evidence as
to similar characters in the primary region of the xylem.
=Coenoxylon.= Zalessky.
_Coenoxylon Scotti_ Zalessky. Prof. Zalessky[776] gave the name
_Coenoxylon_ to a small and incomplete piece of stem of doubtful
provenance but possibly from the Ural Permian beds. The pith, 2 cm.
broad, consists of parenchyma associated in the central region with
numerous large sclerous cells. In one section a sinuous band of
meristematic tissue was observed near the periphery of the pith: the
appearance of this tissue in a photograph given to me by Prof. Zalessky
suggests comparison with occasional strips of similar dividing cells
in the pith of _Lyginopteris_. The secondary wood is composed of
tracheids with 1–2 rows of flattened or hexagonal pits on the radial
walls and narrow uniseriate medullary rays reaching a depth of 15 cells
and with 2–7 oval pits in the field. As in the wood of _Mesopitys
Tchihatcheffi_[777] there are distinct and apparently complete rings of
growth.
It is on the ground of the arrangement and structure of the primary
xylem that Zalessky instituted a new generic name. The primary xylem
forms teeth of variable size which project into the pith from the edge
of the secondary xylem: the prominent portions of the main mass of
primary xylem give off branches, differing considerably in size and
shape, some of which become separated by a comparatively broad band
of parenchyma from the parent xylem-tissue. These bundles anastomose
in their course through the pith and in doing so incorporate between
them patches of parenchyma. The bundles of primary xylem are endarch.
From the centrifugal strands at the periphery of the pith double
leaf-traces are produced which pass almost horizontally through the
secondary wood. As Zalessky points out, the leaf-traces in their dual
nature and in the elongated and narrow form of the tracheal groups,
as seen in tangential section of the secondary wood, bear a close
resemblance to those of _Ginkgo biloba_.
This Russian genus agrees in its double leaf-trace with _Mesoxylon_,
_Mesopitys_, and _Antarcticoxylon_: among recent plants _Ginkgo_ would
seem to be the most closely allied type.
=Parapitys.= Zalessky.
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