Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The designation _Parapitys_[778] has been proposed for a single Upper
Carboniferous species characterised by the possession of secondary wood
like that of _Cordaites_, double leaf-traces, and small mesarch primary
xylem-strands. Nothing is known of the leaves or reproductive organs.
_Parapitys Spenceri_ (Scott).
In 1880 Williamson[779] published a short account of a transverse
section of a specimen found by Mr J. Spencer in Upper Carboniferous
strata near Halifax in Yorkshire which afforded evidence of the
occurrence of double leaf-traces. The following description is taken
from Scott’s account[780] of the species, which he named _Dadoxylon
Spenceri_. The parenchymatous pith, 5–6 mm. in diameter, is obtusely
pentagonal, the prominent angles corresponding to the points of exit
of paired leaf-traces like those of _Mesoxylon_, _Ginkgo_, and other
genera. The secondary xylem consists of narrow tracheids with crowded
multiseriate pits on the radial walls and narrow medullary rays
one-cell broad and 1–8 cells deep. In contact with the inner margin
of the secondary wood are a few small mesarch strands of primary
xylem, the protoxylem and some parenchyma occupying a more or less
central position. A leaf-trace about to enter the secondary xylem is
represented by twin-bundles which retain their double nature as they
traverse the stele, but at a lower level the two components fuse and
appear as single bundles at the outer edge of the pith. The division
of a leaf-trace into two before passing out, as in _Poroxylon_,
constitutes a difference from _Lyginopteris_ in which the division
occurs later.
Zalessky’s generic name _Parapitys_ is an appropriate substitute for
_Dadoxylon_ in view of the presence of separate primary xylem strands,
a feature foreign to typical Dadoxylons which agree with recent
Conifers and Cycads in the absence of vascular strands distinct from
the endarch centrifugal wood. As Scott[781] says, _Parapitys_ ‘is best
regarded as a near ally of _Mesoxylon_.’
=Mesopitys.= Zalessky.
_Mesopitys Tchihatcheffi_ (Goeppert). The genus _Mesopitys_ was
instituted by Zalessky for a Permian species founded by Goeppert[782]
on a piece of decorticated stem from the Kousnetzk basin in Siberia
and afterwards more fully described by Goeppert and Stenzel[783].
The structure of the secondary wood led Goeppert to adopt the name
_Araucarites_ for which Kraus[784] substituted _Araucarioxylon_.
A recent investigation of additional material by Zalessky brought
to light the existence of groups of primary xylem abutting on the
secondary xylem and projecting into the pith, characterised by the
occurrence of spiral protoxylem elements on the inner edge. The
recognition of this important feature justified Zalessky in the
adoption of a new generic term. In general anatomical characters
_Mesopitys_ agrees with _Eristophyton Beinertianum_ (Goepp.) but is
distinguished by the more feebly developed primary-xylem groups and by
their endarch structure.
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