Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The species _Conostoma anglo-germanicum_ agrees closely with _C.
oblongum_ in general form and organisation, but it has eight ribs,
four more prominent than the others, and differs also in other
minor characters from the rather shorter seeds of the type-species.
_Conostoma_ differs from _Lagenostoma_ in the absence of the tubular
prolongation of the lagenostome, the micropyle being like that in
recent Gymnosperms. In _Conostoma_ the tracheid-like elements of the
lateral wall of the lagenostome are a characteristic feature, and no
evidence has been found of the existence of a central core of tissue
such as occupies the centre of the seed-apex in _Lagenostoma_. The
long hairs of _Physostoma_ are represented in _Conostoma_ by the much
smaller mucilaginous cells of the epidermis and in _Lagenostoma_ by
the less closely united mucilage-cells of the superficial layer of the
testa.
=Sphaerostoma.= Benson.
As already pointed out in Chapter +xxix.+ where this genus is described
as probably the seed of _Heterangium_, there is a fairly close general
resemblance between _Sphaerostoma_ and _Lagenostoma_. In the presence
of free apical lobes the former genus resembles _Conostoma_, and
while agreeing with _Lagenostoma_ in its annular pollen-chamber it
is peculiar in the retention of an epidermis over the roof of the
pollen-chamber: as in _Lagenostoma_ the seed is enclosed by an outer
integument or cupule.
=Lagenostoma.= Williamson.
An account of this type of seed is included in the description of
_Lyginopteris_[837]. The more striking peculiarities are exhibited by
the pollen-chamber and the free region of the integument: the annular
pollen-chamber (fig. 493, D, _c_; fig. 409) surrounds a central
nucellar cone and is prolonged upwards as a tube engaging with the
micropyle in contrast to the form of the pollen-chamber and the absence
of a tubular prolongation in _Conostoma_. The tentacles of _Physostoma_
and the short apical lobes of _Conostoma_ are replaced by an apical
cone formed by the coalescence of the integument containing nine
cavities originally filled with parenchyma (figs. 409; 493, B). The
presence of a cupule is a characteristic feature of young seeds, but
from negative evidence in the case of most other seeds it is unsafe
to assume that the cupule of _Lagenostoma_ is an exceptional feature.
The nucellus and testa are united as far as the shoulders of the seed
as in the seeds of recent Cycads and in contrast to their lateral
independence in _Trigonocarpus_, _Stephanospermum_, and other genera.
=Gnetopsis.= Renault.
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