Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This generic name was given by Renault[838] to some small petrified
seeds from the Stephanian of Grand’ Croix and to impressions from the
Commentry coal-field which he believed to belong to some Gnetaceous
plant. Saporta and Marion[839] and other authors have accepted
these seeds as evidence of the existence of Palaeozoic Gnetales: it
has, however, been shown[840] that _Gnetopsis_ has no claim to such
relationship and is a type of seed closely allied to _Conostoma_.
Renault described three species, afterwards adding three from another
locality[841]; the genus is recorded also from Commentry[842] and
Gard[843]. More recently Depape and Carpentier[844] have described
examples from the Westphalian of Valenciennes which they place in
the Pteridosperms in accordance with the conclusion of Oliver and
Salisbury. _Gnetopsis_ has also been discovered by Mr Hemingway in the
Middle Coal Measures of England[845] (fig. 494, H).
_Gnetopsis elliptica_ Renault.
The seeds of this species, slightly oval in section, occur in groups of
2–4 in a cupular investment (fig. 506, E, p. 359) described by Renault
as an ovary but correlated by Oliver and Salisbury with the cupule
which surrounds the single ovule of _Lagenostoma_. The cupule is lined
with hairs similar to those on the wall of the cupule of _Lagenostoma_.
A characteristic feature of the French seeds is the presence of three
or four long plumes of hairs at the apex (fig. 494, E, F). As seen in
fig. 494, E, a small lagenostome (pollen-chamber) rests on the roof of
a broad plinth precisely as in _Conostoma_, and four vascular bundles,
corresponding to the six bundles in _Conostoma_, pass into the apical
cap of sclerous tissue enclosed by a sarcotesta, _sa_ (fig. 494, E,
G). A ‘tent-pole’ prolongation (fig. 494, E, _t_) occurs at the apex
of the prothallus. Renault described a portion of the integument as
consisting of lacunar tissue which Oliver and Salisbury homologise with
the superficial mucilaginous layer of _Conostoma_: this is seen above
the sclerotesta in the apical region of fig. 494, E, _sa_.
_Gnetopsis anglica_ Kidston +MS.+
This species (fig. 494, H) is represented by seeds from the Middle Coal
Measures near Barnsley, Yorkshire, 4 mm. long with apical appendages
at least 2·2 cm. in length and probably four in number. The appendages
do not show the hairs which form a prominent feature in the French
specimens, but this is probably the result of imperfect preservation:
there are indications of hairs on other specimens in Dr Kidston’s
Collection. The type-specimen, in Dr Kidston’s Collection, was
generously lent to me for examination.
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