Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In this genus Brongniart[865] included two species, _Hexapterospermum
stenopterum_ and _H. pachypterum_: the shell is hexagonal in
transverse section, each angle being prolonged as a narrow flange.
In one of the sections figured by Brongniart (fig. 495, E) the
ribs are not fissured: this is said to be a feature distinguishing
_Hexapterospermum_ from _Ptychotesta_, but the occurrence of a
fissured rib in another section suggests that in the structure of
the ribs there is no essential difference between the two genera.
In _Ptychotesta pachypterum_ the testa is prolonged at the chalazal
end as in _Polylophospermum_, and it is possible that there is
no generic difference. Williamson described a cast from the Coal
Measures of Lancashire as _Hexapterospermum_ [= _Hexagonocarpus_]
_Noegerrathi_[866] (fig. 506, H), but in the absence of anatomical
characters it is preferable to avoid the use of Brongniart’s term and
to assign them to Renault’s genus _Hexagonocarpus_[867]. Similarly
the seeds referred by Dr P. Bertrand[868] to _Hexapterospermum_ may
appropriately be included in the genus _Hexagonocarpus_.
[Illustration: Fig. 495. A, _Polypterospermum Renaultii_. B,
_Ptychotesta tenuis_. C, D, _Polylophospermum_ _stephanense_; _sa_,
sarcotesta; _s_, spaces; _v_, vascular bundles. E, _Hexapterospermum
stenopterum_. F, _Diplotesta avellana_. G, _Taxospermum_ _Grüneri_. H,
_Diplotesta Grand’Euryana_. (After Brongniart; C, after Oliver.)]
=Polypterospermum.= Brongniart.
This generic name was proposed for an ovoid seed from St Étienne
described as _Polypterospermum Renaultii_[869]; hexagonal in section
with six deep and sharp flanges at the angles alternating with six
secondary ridges distinguished by their blunter edges and slightly
smaller depth (fig. 495, A). Without further anatomical details it
is hardly possible to say whether or not the species represents a
well-defined generic type, but it is not improbable that a fuller
knowledge would confirm Brongniart’s institution of a new generic
designation. The seeds described by Arber[870] and Kidston[871]
respectively as _Radiospermum ornatum_ and _Polypterospermum ornatum_
are transferred to _Polygonocarpus_ on the ground that they afford no
evidence of anatomical characters of the _Polypterospermum_ type.
=Pachytesta.= Brongniart.
Brongniart[872] established this genus for some unusually large seeds
(fig. 496, A) from the Coal Measures of France reaching a length of
11–12 cm. and resembling in form and size a hen’s egg. Petrified
examples have been described by Brongniart and Renault[873], and
Oliver[874] has given a lucid statement of the more interesting
features of this Permo-Carboniferous genus.
_Pachytesta gigantea_ (Grand’Eury).
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